Publications – Mike Benton's stuff

2024

Mid-Miocene terrestrial carbon isotope shift driven by atmospheric CO2 in the Xining Basin, NE Tibetan Plateau. Palaeogeography, Palaeoclimatology, Palaeoecology 633, 111877 (doi: 10.1016/j.palaeo.2023.111877) (Zhao, Y., Wu, F., Fang, X., Song, Y. and Benton, M.J.). pdf.
Editorial preface to special issue: Recovery of marine ecosystem after the Permian-Triassic mass extinction: New progress from South China. Palaeogeography, Palaeoclimatology, Palaeoecology 633, 111899 (doi: 10.1016/j.palaeo.2023.111899) (Song, H.Y., Hu, S.X.,, Benton, M.J. and Jiang, D.Y.). pdf.
Locomotion and the early Mesozoic success of Archosauromorpha. Royal Society Open Science, 11, 231495. (doi: 10.1098/rsos.231495) (Shipley, A.E., Elsler, A., Singh, S.A., Stubbs, T.L. and Benton, M.J.). pdf.
Predatory synapsid ecomorphology signals growing dynamism of late Palaeozoic terrestrial ecosystems. Communications Biology, 7, 201 (doi: 10.1038/s42003-024-05879-2) (Singh, S.A., Elsler, A., Stubbs, T.L., Rayfield, E.J. and Benton, M.J.). pdf.
Latest Triassic terrestrial microvertebrate assemblages from caves on the Mendip palaeoisland, SW England, at Emborough, Batscombe and Highcroft Quarries. Proceedings of the Geologists’ Association 135, 105-130 (doi: 10.1016/j.pgeola.2023.12.003) (Cawthorne, M., Whiteside, D.I. and Benton, M.J.). pdf.
Finding the world’s oldest mammals: sieving, dialectical materialism, and squabbles. Zoological Journal of the Linnean Society 200, 796–824 (doi: 10.1093/zoolinnean/zlad089) (Benton, M.J., Gill, P.G. and Whiteside, D.I.). pdf.
Universal hyperthermal model for mass extinctions. Journal of Earth Science 35, 704–707 (doi: 10.1007/s12583-024-1985-9). pdf.

The following papers have been published online, but are not paginated yet

The dinosaur boom in the Cretaceous. Special Publications, Geological Society of London 544, online first (doi: 10.1144/SP544-2023-7). pdf.
Onset of the Rhaetian Transgression in deep waters at Lilstock, North Somerset: microvertebrate faunas. Proceedings of the Geologists’ Association online first (doi: 10.1016/j.pgeola.2024.02.001). (Durbin, O.L., Duffin, C.J., Hildebrandt, C. and Benton, M.J.). pdf.

2023

The life of Mary Anning, fossil collector of Lyme Regis: a contemporary biographical memoir by George Roberts. Journal of the Geological Society 180, jgs2022-053 (doi: 10.1144/jgs2022-053) (Taylor, M.A. and Benton, M.J.). pdf.
New information on the cranial anatomy of the Middle Triassic rhynchosaurian reptile Bentonyx sidensis. Acta Palaeontologia Polonica 68, 53-62 (doi: 10.4202/app.01019.2022) (Sethapanichsakul, T., Coram, R.A., and Benton, M.J.). pdf.
The first dinosaurs in China: Dating Late Triassic footprint fossils from the Sichuan Basin. Gondwana Research 117, 261–273 (doi: 10.1016/j.gr.2023.02.003) (Peng, S., Liu, J., Benton, M. J., Jin, X., and Shi, Z.). pdf.
Functional space analyses reveal the function and evolution of the most bizarre theropod manual unguals. Communications Biology 6, 181 (doi: 10.1038/s42003-023-04552-4) (Qin, Z., Liao, C.C., Benton, M.J. and Rayfield, E.J.). pdf. Supplementary data are here.
Comment on “Sexual selection promotes giraffoid head-neck evolution and ecological adaptation”. Science 379, eadd9559 (doi: 10.1126/science.add9559) (Hou, S.K., Shi, Q.Q., Benton, M.J., and Solounias, N.). pdf.
First occurrence of hybodontid teeth in the Luoping Biota (Middle Triassic, Anisian), emphasizing recovery of the marine ecosystem after the end-Permian mass extinction. Palaeogeography, Palaeoclimatology, Palaeoecology, p.111471 (Wen, W., Zhang, Q., Benton, M.J., Kriwet, J., Hu, S., Huang, J., Zhou, C., Cui, X., Ma, Z. and Min, X.). pdf.
The stability and collapse of marine ecosystems during the Permian-Triassic mass extinction. Current Biology 33, 1059-1070. e4 (doi: 10.1016/j.cub.2023.02.007) (Huang, Y.G., Chen, Z.-Q., Roopnarine, P.D., Benton, M.J., Zhao, L.S., Feng, X.Q., and Li. Z.H.). pdf.
Geology and microvertebrate faunas of the Rhaetian Westbury Formation of Doniford Bay, Somerset. Proceedings of the Geologists’ Association 134, 166-182 (doi: 10.1016/j.pgeola.2023.01.004) (Tayler, J., Duffin, C.J., Hildebrandt, C., Parker, A., and Benton, M.J.). pdf. Supplementary materials are here.
The locomotor ecomorphology of Mesozoic marine reptiles. Palaeontology 66, e12645 (doi: 10.1111/pala.12645) (Gutarra, S., Stubbs, T.L., Moon, B.C., Heighton, B.H., and Benton, M.J.). pdf. Supplementary information is here. Voted the best paper in Palaeontology for 2023.
Palaeobiology: Rapid succession during mass extinction. Current Biology 33, R429-R452 (doi: 10.1016/j.cub.2023.04.043). pdf.
Unique dentition of rhynchosaurs and their two-phase success as herbivores in the Triassic. Palaeontology 66, e12654 (doi: 10.1111/pala.12654) (Sethapanichsakul, T., Coram, R.A. and Benton, M.J.). pdf. Supplementary information is in MorphoSource.
Extended embryo retention and viviparity in the first amniotes. Nature Ecology & Evolution 7, 1131-1140 (doi: 0.1038/s41559-023-02074-0) (Jiang, B., He, Y., Elsler, A., Wang, S., Keating, J.N., Song, J., Kearns, S.L. and Benton, M.J.). pdf.
Fungal-induced fossil biomineralization. Current Biology 33, 2417-2424 (doi: 10.1016/j.cub.2023.04.067) (Luo, M., Li, Z., Su, M., Gadd, G.M., Yin, Z., Benton, M.J., Pan, Y., Zheng, D., Zhao, T., Li, Z. and Chen, Y.). pdf.
Permian–Middle Triassic floral succession in North China and implications for the great transition of continental ecosystems. Bulletin of the Geological Society of America 135, 1747–1767 (doi: 10.1130/B36316.1) (Shu, W.C., Tong, J.N., Yu, J.X., Hilton, J., Benton, M.J., Shi, X., Diez, J.B., Wignall, P.B., Chu, D.L., Tian, L., Yi, Z.X., and Mao, Y.D.). pdf. Supplementary materials are here.
Volcanic geological sites in UGGp European geoparks: special issue. Geoconservation Research 6, i-x (doi: 10.30486/gcr.2023.1990600.1143) (Nunes, J.C. and Benton, M.J.). pdf.
The curious case of Central Park’s dinosaurs: The destruction of Benjamin Waterhouse Hawkins’ Paleozoic Museum revisited. Proceedings of the Geologists’ Association 134, 344-360 (doi: 10.1016/j.pgeola.2023.04.004) (Coules, V.J. and Benton, M.J.). pdf.
Allometric wing growth links parental care to pterosaur giantism. Proceedings of the Royal Society of London B 290, 20231102 (doi: 0.1098/rspb.2023.1102) (Yang, Z., Jiang, B., Benton, M.J., Xu, X., McNamara, M.E. and Hone, D.W.E.). pdf.
First filter feeding in the Early Triassic: cranial morphological convergence between Hupehsuchus and baleen whales. BMC Ecology and Evolution 23, 36 (doi: 10.1186/s12862-023-02143-9) (Fang, Z.C., Li, J.L., Yan, C.B., Zou, Y.R., Tian, L., Zhao, B., Benton, M.J., Cheng, L., and Lai, X.L.). pdf.
Rapid neck elongation in Sauropterygia (Reptilia: Diapsida) revealed by a new basal pachypleurosaur from the Lower Triassic of China. BMC Ecology and Evolution 23, 44 (doi: 10.1186/s12862-023-02150-w) (Liu, Q.L., Cheng, L., Stubbs, T.L., Moon, B.C., Benton, M.J., Yan, C.B. and Tian, L.). pdf.
Bayesian analyses indicate bivalves did not drive the downfall of brachiopods following the Permian-Triassic mass extinction. Nature Communications 14, 5566 (doi: 10.1038/s41467-023-41358-8) (Guo, Z., Flannery-Sutherland, J.T., Benton, M.J. and Chen, Z.Q.). pdf.
Divergent strategies in cranial biomechanics and feeding ecology of the ankylosaurian dinosaurs. Scientific Reports 13, 18242 (doi: 10.1038/s41598-023-45444-1) (Ballell, A., Mai, B. and Benton, M.J.). pdf.
Crustacean microcoprolites from the Middle Triassic Luoping Biota, China: Evidence for primary producers in the first Modern-type marine ecosystems. Palaeogeography, Palaeoclimatology, Palaeoecology 632, 111840 (doi: 10.1016/j.palaeo.2023.111840) (Chen, Y., Luo, M., Benton, M.J., Hu, S., Zhang, Q., Ji, C., Huang, J., Kou, Q., Zhou, C. and Wen, W.). pdf.
A Coelophysoidea (Dinosauria, Theropoda) femur from the Tytherington fissures (Rhaetian, Late Triassic), Bristol, UK. Proceedings of the Geologists’ Association 134, 562-572 (doi: 10.1016/j.pgeola.2023.07.005). (Kirmse, J.P., Benton, M.J., Hildebrandt, C., Langer, M.C. and Marsola, J.C.). pdf.
Exceptionally preserved conodont natural assemblages from the Middle Triassic Luoping Biota, Yunnan Province, China: implications for architecture of conodont feeding apparatus. Journal of Earth Science 34, 1762-1776 (doi: 10.1007/s12583-022-1793-z). (Huang, J., Martínez-Pérez, C., Zhang, Q., Zhang, K., Luo, M., Wen, W., Zhou, C., Min, X., Ma, Z., Xie, T., Benton, M.J. and Donoghue, P.C.J.). pdf.

2022

Phylogenetic classification and evolution of Early Triassic conodonts. Palaeogeography, Palaeoclimatology, Palaeoecology 585, 110731 (doi: 10.1016/j.palaeo.2021.110731) (Bai, R.Y., Song, H.J., Benton, M.J., and Tian, L.). pdf. Supplementary materials are available here.
The naming of the Permian System. Journal of the Geological Society 179, jgs2021-037 (doi: 10.1144/jgs2021-037) (Benton, M.J. and Sennikov, A.G.) pdf. Supplementary material (key Russian papers and translations), available here.
Walking with early dinosaurs: appendicular myology of the Late Triassic sauropodomorph Thecodontosaurus antiquus. Royal Society Open Science 9, 211356 (doi: 10.1098/rsos.211356) (Ballell, A., Rayfield, E.J., and Benton, M.J.). pdf. Supplementary materials are available here.
The Angiosperm Terrestrial Revolution and the origins of modern biodiversity. New Phytologist 233, 2017–2035 (Benton, M.J., Wilf, P., and Sauquet, H.S.). pdf. One of the top-ten most read reviews in New Phytologist for 2021 and 2022.
Slow and fast evolutionary rates in the history of lepidosaurs. Palaeontology 65, e12579 (doi: 10.1111/pala.12579) (Herrera-Flores, J.A., Elsler, A., Stubbs, T.L., and Benton, M.J.). pdf. Supplementary materials are available here and at Dryad. Read a great report at Scientific American by Riley Black.
Reply to: ‘Reconstructed evolutionary patterns from crocodile-line archosaurs demonstrate the impact of failure to log-transform body size data.’ Communications Biology 5, 170 (doi: 10.1038/s42003-022-03072-x) (Stockdale, M.T. and Benton, M.J.). pdf.
Phylogenetic relationships of the European trilophosaurids Tricuspisaurus thomasi and Variodens inopinatus. Journal of Vertebrate Paleontology 41, e1999250 (doi: 10.1080/02724634.2021.1999250) (Chambi-Trowell, S.A.V., Whiteside, D.I., Skinner, M., Benton, M.J., and Rayfield, E.J.). pdf. Supplemental data are available here.
Climate, competition, and the rise of mosasauroid ecomorphological disparity. Palaeontology 65, e12590 (doi: 10.1111/pala.12590) (Cross, S.R.R., Moon, B.C., Stubbs, T.L., Rayfield, E.J., and Benton, M.J.). pdf. Supplemental data are available here.
Large size in aquatic tetrapods compensates for high drag caused by extreme body proportions. Communications Biology 5, 380 (doi: https://doi.org/10.1038/s42003-022-03322-y) (Gutarra, S. Stubbs, T.L., Moon, B.C., Palmer, C., and Benton, M.J.). pdf. Supplemental data are available here.
Allometric analysis sheds light on the systematics and ontogeny of anurognathid pterosaurs. Journal of Vertebrate Paleontology 41, e2028796 (doi: 10.1080/02724634.2021.2028796) (Yang, Z.X., Benton, M.J., Hone, D.W.E., Xu, X., McNamara, M.E., and Jiang, B.Y.). pdf. Supplemental data are available here.
A colourful view of the origin of pterosaur feathers. Nature 604, 630-631 (doi: https://doi.org/10.1038/d41586-022-01036-z). pdf.
The Jurassic rise of squamates as supported by lepidosaur disparity and evolutionary rates. eLife 11, e66511. (doi: 10.7554/eLife.66511) (Bolet, A., Stubbs, T.L., Herrera-Flores, J.A., and Benton, M.J.). pdf. Supplemental data are here.
Microvertebrates from the Rhaetian bone beds at Westbury Garden Cliff, near Gloucester, UK. Proceedings of the Geologists’ Association 133, 119-136 (doi: 10.1016/j.pgeola.2022.01.002) (Williams, H., Duffin, C.J., Hildebrandt, C., Parker, A., Hutchinson, D., and Benton, M.J.). pdf. Supplemental data are available here.
Global diversity dynamics in the fossil record are regionally heterogeneous. Nature Communications 13, 2751 (doi: 10.1038/s41467-022-30507-0) (Flannery Sutherland, J.T., Silvestro, D., and Benton, M.J.). pdf. Supplemental data are available here.
Triassic revolution. Frontiers in Earth Science 10, 899541 (doi: 10.3389/feart.2022.899541) (Benton, M.J. and Wu, F.X.). pdf.
Resilience of infaunal ecosystems during the Early Triassic greenhouse Earth. Science Advances 8, eabo0597 (doi: 10.1126/sciadv.abo059) (Feng, X.Q., Chen, Z.Q., Benton, M.J., Su, C.M., Bottjer, D.J., Cribb, A.T., Li, Z.H., Zhao, L.S., Zhu, G.Y., Huang, Y.G., and Guo, Z). pdf. Supplementary materials are here.
Post-extinction recovery of the Phanerozoic oceans and biodiversity hotspots. Nature 607, 507–511 (doi: 10.1038/s41586-022-04932-6) (Cermeño, P., García-Comas, C., Pohl, A., Williams, S., Benton, M.J., Chaudhary, C., Le Gland, G., Müller, D., Ridgwell, A., and Vallina, S.M.). pdf. Supplementary information is here. History of the oceans and biodiversity in 26 seconds is here.
A new horseshoe crab from the Permian-Triassic transition of South China: Xiphosurids as key components of post-extinction trophic webs. Palaeogeography, Palaeoclimatology, Palaeoecology 602, 111178 (Hu, S.X., Feldmann, R.M., Schweitzer, C.E., Benton, M.J., Huang, J.Y., Wen, W., Min, X., Zhang, Q.Y., Zhou, C.Y., and Ma, Z.X.). pdf. Supplementary materials are here.
Improving paleoenvironment in North China aided Triassic biotic recovery on land following the end-Permian mass extinction. Global and Planetary Change 216, 103914 (Zhu, Z.C., Liu, Y.Q., Kuang, H.W., Newell, A.J., Peng, N., Cui, M.M., and Benton, M.J.). pdf. Supplementary materials are here.
Reply to: Revisiting life history and morphological proxies for early mammaliaform metabolic rates. Nature Communications 13, 5564 (doi: 10.1038/s41467-022-32716-z) (Newham, E., Gill, P.G., Brewer, P., Benton, M.J., Fernandez, V., Gostling, N.J., Habertür, D., Jernvall, J., Kankaanpää, T., Kallonen, A., Navarro, C., Pacureanu, A., Richards, K., Robson Brown, K., Schneider, P., Suhonen, H., Tafforeau, P., Williams, K.A., Zeller-Plumhoff, B., and Corfe, I.J.). pdf. Supplementary files here.
Impact of the Jenkyns Event (early Toarcian) on dinosaurs: Comparison with the Triassic/Jurassic transition. Earth-Science Reviews 234, 104196 (doi: 10.1016/j.earscirev.2022.104196) (Reolid, M., Ruebsam, W., and Benton, M.J.). pdf. Supplemental files are here.
Hybodontiform shark remains (Chondrichthyes, Elasmobranchii) from the Lower Triassic of Yunnan Province, China, with comments on hybodontiform diversity across the PTB. Journal of Vertebrate Paleontology 42, e2108712 (doi: 10.1080/02724634.2022.2108712) (Wen, W., Kriwet, J., Zhang, Q.Y., Benton, M.J., Duffin, C.J., Huang, J.Y., Zhou, C.Y., Hu, S.X., and Ma, Z.X.). pdf.
Dietary niche partitioning in Early Jurassic ichthyosaurs from Strawberry Bank. Journal of Anatomy 241, 1409-1423 (doi: 10.1111/joa.13744) (Jamison-Todd, S., Moon, B.C., Rowe, A.J., Williams, M., and Benton., M.J.). pdf. Supplementary files are here. Featured on the front cover for December 2022.
A new exposure of the North Curry Sandstone Member (Dunscombe Mudstone Formation, Mercia Mudstone Group: Carnian, Triassic), near Taunton, Somerset (UK): The location of Charles Moore’s vertebrate specimens resolved. Proceedings of the Geologists’ Association 133, 526-537 (doi: 10.1016/j.pgeola.2022.06.009) (Dawson, G.J., Burley, S.D., Ruffell, A., Benton, M.J., and Duffin, C.J.). pdf.
A Triassic crown squamate. Science Advances 8, eabq8274 (doi: 10.1126/sciadv.abq8274) (Whiteside, D.I., Chambi-Trowell, A.A.V., and Benton, M.J.). pdf. Supplementary materials are here. Three-dimensional images and phylogenetic analysis code at Dryad.
Dental form and function in the early feeding diversification of dinosaurs. Science Advances 8, eabq5201 (doi: 10.1126/sciadv.eabq5201) (Ballell, A., Benton, M.J., and Rayfield, E.J.). pdf. Supplementary materials are here.
Establishment of the Luoping Biota National Geopark in Yunnan, China. Geoconservation Research 5, 261-284 (doi: 10.30486/gcr.2022.1962749.1111) (Benton, M.J., Hu, S.X., Zhang, Q.Y., Xie, T., Zhou, C.Y., and Huang, J.Y.). pdf.
Dinosaur extinctions related to the Jenkyns Event (early Toarcian, Jurassic). Spanish Journal of Palaeontology 37, 123-140 (doi: 10.7203/sjp.25683) (Reolid, M., Ruebsam, W., and Benton, M.J.). pdf.

2021

Environmental drivers of body size evolution in crocodile-line archosaurs. Communications Biology 4, 38 (doi: 10.1038/s42003-020-01561-5) (Stockdale, M.T. and Benton, M.J). pdf. Supplementary data are here.
Evolution of ecospace occupancy by Mesozoic marine tetrapods. Palaeontology 64, 31–49 (doi: 10.1111/pala.12508) (Reeves, J.C., Moon, B.C., Benton, M.J., and Stubbs, T.L.). pdf. Supplementary data in Dryad.
Reply to Walkden, Fraser and Simms (2021): The age and formation mechanisms of Late Triassic fissure deposits, Gloucestershire, England: Comments on Mussini, G., Whiteside, D. I., Hildebrandt C. and Benton M.J. Proceedings of the Geologists’ Association 132, 138-141 (doi: 10.1016/j.pgeola.2020.12.001) (Whiteside, D.I. and Benton, M.J.). pdf. Download supplementary information here.
Ecomorphological diversification of squamates in the Cretaceous. Royal Society Open Science 8, art. 201961 (doi: 10.1098/rsos.201961) (Herrera-Flores, J.A., Stubbs, T.L., and Benton, M.J. pdf. Download data here.
Ecological dynamics of terrestrial and freshwater ecosystems across three mid-Phanerozoic mass extinctions from northwest China. Proceedings of the Royal Society B 288, 20210148 (doi: 10.1098/rspb.2021.0148) (Huang, Y.G., Chen, Z.Q., Roopnarine, P.D., Benton, M.J., Yang, W., Liu, J., Zhao, L.S. and Guo, Z.). pdf. Supplementary data are available here.
Ecological opportunity and the rise and fall of crocodylomorph evolutionary innovation. Proceedings of the Royal Society B 288, 20210069 (doi: 10.1098/rspb.2021.0069) (Stubbs, T.L., Pierce, S.E., Elsler, A., Anderson, P.S.L., Rayfield, E.J., and Benton, M.J.) pdf. Supplementary data are available here.
Microvertebrates from the Rhaetian basal bone bed of Saltford, near Bath, SW England. Proceedings of the Geologists’ Association 132, 174-187 (doi: 10.1016/j.pgeola.2020.11.003) (Moreau, M., Duffin, C.J., Hildebrandt, C., Hutchinson, D., Parker, A., Carpenter, S., and Benton, M.J.). pdf.
Testing for a dietary shift in the Early Cretaceous ceratopsian dinosaur Psittacosaurus lujiatunensis. Palaeontology 64, 371-384 (doi: 10.1111/pala.12529) (Landi, D., King, L., Zhao, Q., Rayfield, E.J., and Benton, M.J.). pdf. Data available from the Dryad Digital Repository.
Testing the relationship between marine transgression and evolving island palaeogeography using 3D GIS: an example from the Late Triassic of SW England. Journal of the Geological Society 178, jgs2020-158 (doi: 10.1144/jgs2020-158) (Lovegrove, J., Newell, A.J., Whiteside, D.I., and Benton, M.J.). pdf. Data are available here.
Niche partitioning shaped herbivore macroevolution through the early Mesozoic. Nature Communications 12, art. 2796 (doi: 10.1038/s41467-021-23169-x) (Singh, S.A., Elsler, A., Stubbs, T.L., Bond, R., Rayfield, E.R., and Benton, M.J.). pdf. Supplementary data here and here.
Dinosaur biodiversity declined well before the asteroid impact, influenced by ecological and environmental pressures. Nature Communications 12, art. 3833 (doi: 10.1038/s41467-021-23754-0) (Condamine, F.L., Guinot, G., Benton, M.J. and Currie, P.J.). pdf. Supplementary data here.
Ontogenetic endocranial shape change in alligators and ostriches and implications for the development of the nonavian dinosaur endocranium. The Anatomical Record 304, 1759-1775 (doi: 10.1002/ar.24579) (Hu, K., King, J.L., Romick, C.A., Dufeau, D.L., Witmer, L.M., Stubbs, T.L., Rayfield, E.R, and Benton, M.J.). pdf. Supplementary figure.
Decoupling of morphological disparity and taxonomic diversity during the end-Permian mass extinction. Paleobiology 47, 402-417 (doi: 10.1017/pab.2020.57). (Wan, J.Y., Foster, W.J., Tian, L., Stubbs, T.L., Benton, M.J., Qiu, X.C., and Yuan, A.H.). pdf. Data available from the Dryad Digital Repository.
European UNESCO geoparks: introduction to Part 1. Geoconservation Research 4, 1-5 (doi: 10.30486/gcr.2021.683780). pdf.
The North West Highlands UNESCO Global Geopark: oldest fossils in Europe. Geoconservation Research 4, 25-42 (doi: 10.30486/gcr.2020.1911147.1034) (Benton, M.J., Brasier, A., Harrison, P., and Hamlet, L.). pdf.
Growth and miniaturization among alvarezsauroid dinosaurs. Current Biology 31, 3687–3693 (doi: 10.1016/j.cub.2021.06.013) (Qin, Z.C., Qi, Z., Choiniere, J.N., Clark, J.M., Benton, M.J. and Xu, X.). pdf.
Strong support for a heterogeneous speciation decline model in Dinosauria: a response to claims made by Bonsor et al. (2020). Royal Society Open Science 8, 202143 (doi: 10.1098/rsos.202143) (Sakamoto, M., Benton, M.J., and Venditti, C.). pdf.
Triassic tragedy—a bone bed in the Otter Sandstone of East Devon, south-west England. Geology Today 37, 176-183 (doi: 10.1111/gto.12367) (Coram, R.A., Radley, J.D. and Benton, M.J.). pdf.
The braincase, brain and palaeobiology of the basal sauropodomorph dinosaur Thecodontosaurus antiquus. Zoological Journal of the Linnean Society 193, 541-562 (doi: 10.1093/zoolinnean/zlaa157) (Ballell, A., King, J. L., Neenan, J. M., Rayfield, E. J., and Benton, M.J.). pdf. Supplementary data.
Plant resilience and extinctions through the Permian to Middle Triassic on the North China Block: A multilevel diversity analysis of macrofossil records. Earth-Science Reviews 223, 103846 (doi: 10.1016/j.earscirev.2021.103846) (Xiong, C.H., Wang, J.S., Huang, P., Cascales-Miñana, B., Cleal, C.J., Benton, M.J., and Xue, J.Z.). pdf. Supplementary data.
The diversity of Triassic South American sphenodontians: a new basal form, clevosaurs, and a revision of rhynchocephalian phylogeny. Journal of Systematic Palaeontology 19, 787-820 (doi: 10.1080/14772019.2021.1976292) (Chambi-Trowell, A.A.V., Martinelli, A.G., Whiteside, D.I., de Vivar, P.R.R., Soares, M.B., Schultz, C.L., Gill, P.G., Benton, M.J., and Rayfield, E.J.). pdf. Supplementary data.
The origin of endothermy in synapsids and archosaurs and arms races in the Triassic. Gondwana Research 100, 261-289 (Benton, M.J.) (doi: 10.1016/j.gr.2020.08.003). pdf.
A thing with feathers. Current Biology 31, R1403-R1409 (doi: 10.1016/j.cub.2021.09.064) (Benton, M.J., Currie, P.J., and Xu, X.). pdf.
An injured pachypleurosaur (Diapsida: Sauropterygia) from the Middle Triassic Luoping Biota indicating predation pressure in the Mesozoic. Scientific Reports 11, 21818 (doi: 10.1038/s41598-021-01309-z) (Liu, Q.L., Yang, T.L., Cheng, L., Benton, M.J., Moon, B.C., Yan, C.B., An, Z.H., and Tian, L.). pdf.
Biostratigraphic significance and geometric morphometrics of Euestheria gutta (Crustacea: Branchiopoda: Spinicaudata): An index fossil of continental Permian–Triassic transitional beds. Geological Journal 56, 6176-6188 (doi: 10.1002/gj.4096) (Miao, X., Chu, D.L., Tong, J.N., Benton, M.J., Yu, J.X., Luo, L., Shu, W.C., and Wu, Y.Y.). pdf.
European UNESCO geoparks: introduction to Part II. Geoconservation Research 4, 290-298 (doi: 10.30486/gcr.2021.687062). pdf.

2020

Anatomy of a Late Triassic Bristol fissure: Tytherington fissure 2. Proceedings of the Geologists’ Association 131, 73-93 (doi: 10.1016/j.pgeola.2019.12.001) (Mussini, G., Whiteside, D.I., Hildebrandt, C. and Benton, M.J.). pdf. Supplementary file 1. Supplementary file 2.
Morphological disparity in theropod jaws: comparing discrete characters and geometric morphometrics. Palaeontology 63, 283-299 (doi: 10.1111/pala.12455) (Schaeffer, J., Benton, M.J., Rayfield, E.J. and Stubbs, T. L.). pdf. Data for this study are available in the Dryad Digital Repository.
Variable preservation potential and richness in the fossil record of vertebrates. Palaeontology 63, 313-329 (doi: 10.1111/pala.12458) (Walker, F.M., Dunhill, A.M. and Benton, M.J.). pdf. Data for this study are available in the Dryad Digital Repository.
The oldest lambeosaurine dinosaur from Europe: insights into the arrival of Tsintaosaurini. Cretaceous Research 107, 104286 (doi: 10.1016/j.cretres.2019.104286) (Conti, S., Vila, B., Sellés, A.G., Galobart, À., Benton, M.J., Prieto-Márquez, A.). pdf. Supplementary data as a zip file.
The impact of the pull of the recent on extant elasmobranchs. Palaeontology 63, 369-374 (doi: 10.1111/pala.12478) (Pimiento, C. and Benton, M.J.). pdf. Data for this study are available in the Dryad Digital Repository.
Three-dimensional tomographic study of dermal armour from the tail of the Triassic aetosaur Stagonolepis robertsoni. Scottish Journal of Geology 56, 55-62 (doi: 10.1144/sjg2019-026) (Keeble, E. and Benton, M.J.). pdf. Supplementary material (rotating 3D images) available here.
Experimental investigation of insect deposition in lentic environments and implications for formation of konservat lagerstätten. Palaeontology 63, 565-578 (doi: 10.1111/pala.12472) (Tian, Q.Y., Wang, S.Y., Yang, Z.X., McNamara, M.E., Benton, M.J. and Jiang, B.Y.). pdf. Data for this study are available in the Dryad Digital Repository.
Predicting biotic responses to future climate warming with classic ecogeographic rules. Current Biology 30, R737-R758 (doi: 10.1016/j.cub.2020.06.003) (Tian, L. and Benton, M.J.). pdf. News story in Science is here.
Intensifying aeolian activity following the end Permian mass extinction: Evidence from the Late Permian–Early Triassic terrestrial sedimentary record of the Ordos Basin, North China. Sedimentology 67, 2691-2720 (doi: 10.1111/sed.12716) (Zhu, Z.C., Kuang, H.W., Liu, Y.Q., Benton, M.J., Newell, A.J., Xu, H., An, W., Ji, A.A., Xu, S.C., Neng, N., and Zhai, Q.G). pdf.
Footprints of marine reptiles from the Middle Triassic (Anisian-Ladinian) Guanling Formation of Guizhou Province, southwestern China: The earliest evidence of synchronous style of swimming. Palaeogeography, Palaeoclimatology, Palaeoecology 558, 109943 (doi: 10.1016/j.palaeo.2020.109943) (Xing, L.D., Klein, H., Lockley, M.G., Wu, X.C., Benton, M.J., Zeng, R., and Romilio, A.). pdf.
Migration controls extinction and survival patterns of foraminifers during the Permian-Triassic crisis in South China. Earth-Science Reviews 209, 103329 (doi: 10.1016/j.earscirev.2020.103329) (Liu, X.K., Song, H.J., Bond, D.G.P., Tong, J.N., and Benton, M.J.). pdf.
Body dimensions of the extinct giant shark Otodus megalodon: a 2D reconstruction. Scientific Reports 10, 14596 (doi: 10.1038/s41598-020-71387-y) (Cooper, J.A., Pimiento, C., Ferrón, H.G., and Benton, M.J.). pdf. Supplement. Data are available here.
Extinction and dawn of the modern world in the Carnian (Late Triassic). Science Advances 6, eaba0099 (doi: 10.1126/sciadv.aba0099) (Dal Corso, J., Bernardi, M., Sun, Y.D., Song, H.J., Seyfullah, L.J., Preto, N., Gianolla, P., Ruffell, A., Kustascher, E., Roghi, G., Merico, A., Hohn, S., Schmidt, A.R., Marzoli, A., Newton, R.J., Wignall, P.B., and Benton, M.J.) pdf. Read popular account at The Conversation.
Verifiability of genus-level classification under quantification and parsimony theories: a case study of follicucullid radiolarians. Paleobiology 46, 337-355 (doi: 10.1017/pab.2020.28) (Xiao, Y.F., Suzuki, N., He, W.H., Benton, M.J., Yang, T.L., and Cai, C.Y.). pdf.
Osteological redescription of the Late Triassic sauropodomorph dinosaur Thecodontosaurus antiquus based on new material from Tytherington, southwestern England. Journal of Vertebrate Paleontology 40, e1770774 (doi: 10.1080/02724634.2020.1770774) (Ballell, A., Rayfield, E.J. and Benton, M.J.). pdf. Download Supplement.
Reptile-like physiology in Early Jurassic stem-mammals. Nature Communications 11, 5121 (doi: 10.1038/s41467-020-18898-4) (Newham, E., Gill, P.G., Brewer, P., Benton, M.J., Fernandez, V., Gostling, N.J., Habertür, D., Jernvall, J., Kankaanpää, T., Kallonen, A., Navarro, C., Pacureanu, A., Richards, K., Robson Brown, K., Schneider, P., Suhonen, H., Tafforeau, P., Williams, K.A., Zeller-Plumhoff, B., and Corfe, I.J.). pdf. Supplementary files here.
Early Triassic terrestrial tetrapod fauna: a review. Earth-Science Reviews 210, 103331 (doi: 10.1016/j.earscirev.2020.103331) (Romano, M., Bernardi, M., Petti, F.M., Rubidge, B., Hancox, J., and Benton, M.J.). pdf.
Beginning of Mesozoic marine overstep of the Mendips: The Rhaetian and its fauna at Hapsford Bridge, Vallis Vale, Somerset, UK. Proceedings of the Geologists’ Association 131, 535-561 (doi: 10.1016/j.pgeola.2020.02.005) (Ronan, J., Duffin, C.J., Hildebrandt, C., Parker, A., Hutchinson, D., Copp, C., and Benton, M.J.). pdf. Excel file of data.
Late Triassic island dwarfs? Terrestrial tetrapods of the Ruthin fissure (South Wales, UK) including a new genus of procolophonid. Proceedings of the Geologists’ Association 131, 578-594 (doi: 10.1016/j.pgeola.2020.04.005) (Skinner, M., Whiteside, D.I., and Benton, M.J.). pdf. Download Excel file of data.
150 million years of sustained increase in pterosaur flight efficiency. Nature 587, 83-86 (doi: 10.1038/s41586-020-2858-8) (Venditti, C., Baker, J., Benton, M.J., Meade, A., and Humphries, S.). pdf. Article in The Conversation.
Geological control on dinosaurs’ rise to dominance: Late Triassic ecosystem stress by relative sea level change. Terra Nova 32, 434–441 (doi: 10.1111/ter.12480) (Klausen, T.G., Paterson, N.W. and Benton, M.J.). pdf.
An effect size statistical framework for investigating sexual dimorphism in non-avian dinosaurs and other extinct taxa, Biological Journal of the Linnean Society 131, 231–273 (doi: 10.1093/biolinnean/blaa105) (Saitta, E.T., Stockdale, M.T., Longrich, N.R., Bonhomme, V., Benton, M.J., Cuthill, I.C., and Makovicky, P.J.). pdf. Supplement.
Geoconservation in the Carpathian area: Special Issue. Geoconservation Research 3(2), 1-7 (Benton, M.J., Nemeth, K., Makovic, S.B., and Vasiljevic, D.A. (doi: 10.30486/gcr.2020.1913822.1058). pdf.
Biomechanical properties of the jaws of two species of Clevosaurus and a reanalysis of rhynchocephalian dentary morphospace. Palaeontology 63, 919–939 (doi: 10.1111/pala.12493) (Chambi-Trowell, S.A.V., Whiteside, D.I., Rayfield, E.R., and Benton, M.J.). pdf. Additional data for this study are available in the Dryad Digital Repository.
An enigmatic neodiapsid reptile from the Middle Triassic of England. Journal of Vertebrate Paleontology 40, e1781143, online first (doi: 10.1080/02724634.2020.1781143) (Cavicchini, I., Zaher, M., and Benton, M.J.). pdf. Supplement.
Reply to: No protofeathers on pterosaurs. Nature Ecology & Evolution 4, 1592-1593 (doi: 10.1038/s41559-020-01309-8) (Yang, Z.X., Jiang, B.Y., McNamara, M.E., Kearns, S.L., Pittman, M., Kaye, T.G., Orr, P.J., Xu, X., and Benton, M.J.). pdf.
Response to Delhey et al. Current Biology 30, R1408 (doi: 10.1016/j.cub.2020.10.073) (Tian, L. and Benton, M.J.). pdf. News story [Kean, S. 2021. Will warming make animals darker–or lighter? Science 371, 115] is here.
Fish and crab coprolites from the latest Triassic of the UK: From Buckland to the Mesozoic Marine Revolution. Proceedings of the Geologists’ Association 131, 699-721 (doi: 10.1016/j.pgeola.2020.07.011) (Cueille, M., Green, E., Duffin, C.J., Hildebrandt, C., and Benton, M.J.). pdf. Download Excel file of data.

2019

Pterosaur integumentary structures with complex feather-like branching. Nature Ecology & Evolution 3, 24-30 (doi: 10.1038/s41559-018-0728-7) (Yang, Z.X., Jiang, B.Y., McNamara, M.E., Kearns, S.L., Pittman, M., Kaye, T.G., Orr, P.J., Xu, X., and Benton, M.J.) pdf. Supplementary file. News & views commentary.
The mosasaur fossil record through the lens of fossil completeness. Palaeontology 62, 51-75 (doi: 10.1111/pala.12381). (Driscoll, D.A., Dunhill, A.M., Stubbs, T.L. and Benton, M.J.). pdf. Data for this study are available in the Dryad Digital Repository.
Palaeoenvironmental reconstruction and biostratinomic analysis of the Jurassic Yanliao Lagerstätte in northeastern China. Palaeogeography, Palaeoclimatology, Palaeoecology 514, 739-753 (doi: 10.1016/j.palaeo.2018.09.030) (Yang, Z., Wang, S.Y., Tian, Q.Y., Wang, B., Hethke, M., McNamara, M.E., Benton, M.J., Xu, X. and Jiang, B.Y.). pdf.
Early Middle Triassic trace fossils from the Luoping Biota, southwestern China: Evidence of recovery from mass extinction. Palaeogeography, Palaeoclimatology, Palaeoecology 515, 6-22 (doi: 10.1016/j.palaeo.2017.11.028) (Luo, M., Shi, G.R., Hu, S.X., Benton, M.J., Chen, Z.-Q., Huang, J.Y., Zhang, Q.Y., Zhou, C.Y., and Wen, W.). pdf.
Archosauromorph extinction selectivity during the Triassic-Jurassic mass extinction. Palaeontology 62, 211-224 (doi: 10.1111/pala.12399) (Allen, B.J., Stubbs, T.L., Benton, M.J. and Puttick, M.N.). pdf. Supplementary file, at Dryad. Read a Scientific American blog here by Brian Switek.
Reply to comments on: Macroevolutionary patterns in Rhynchocephalia: is the tuatara (Sphenodon punctatus) a living fossil? Palaeontology 62, 335-338 (doi: 10.1111/pala.12404) (Herrera-Flores, J., Stubbs, T.L., and Benton, M.J.). pdf.
The Middle Triassic procolophonid Kapes bentoni: computed tomography of the skull and skeleton. Papers in Palaeontology 5, 111-138 (doi: 10.1002/spp2.1232) (Zaher, M., Coram, R.A., and Benton, M.J.). pdf.
Does exceptional preservation distort our view of disparity in the fossil record? Proceedings of the Royal Society B 286, 20190091 (doi: 10.1098/rspb.2019.0091) (Flannery Sutherland, J., Moon, B.C., Stubbs, T.L., and Benton, M.J.). pdf. Supplementary file.
Environmental instability prior to end-Permian mass extinction reflected in biotic and facies changes on shallow carbonate platforms of the Nanpanjiang Basin (South China). Palaeogeography, Palaeoclimatology, Palaeoecology 519, 23-36 (doi: 10.1016/j.palaeo.2018.05.011) (Tian, L., Tong, J.N., Xiao, Y.F., Benton, M.J., Song, H.Y., Song, H.J., Liang, L., Wu, K., Chu, D.L., and Algeo, T.J.). pdf.
Mixed continental-marine biotas following the Permian-Triassic mass extinction in South and North China. Palaeogeography, Palaeoclimatology, Palaeoecology 519, 95-107 (doi: 10.1016/j.palaeo.2017.10.028) (Chu, D.L., Tong, J.N., Benton, M.J., Yu, J.X., and Huang, Y.F.). pdf.
A diverse trackway-dominated marine ichnoassemblage from the Lower Triassic in the northern Paleotethys: Ichnology and implications for biotic recovery. Palaeogeography, Palaeoclimatology, Palaeoecology 519, 124-140 (doi: 10.1016/j.palaeo.2017.11.059) (Feng, X.Q., Chen, Z.-Q., Benton, M.J., Wu, S.Q., Bottjer, D.J., and Thompson, J.R.). pdf.
Effects of body plan evolution on the hydrodynamic drag and energy requirements of swimming in ichthyosaurs. Proceedings of the Royal Society B 286: 20182786 (doi: 10.1098/rspb.2018.2786) (Gutarra, S., Moon, B.C., Rahman, I.A., Palmer, C., Lautenschlager, S., Brimacombe, A.J., and Benton, M.J.). pdf. Supplementary file. Supplementary data.
Diversity in rhynchocephalian Clevosaurus skulls based on CT reconstruction of C. hudsoni and C. cambrica. Acta Palaeontologia Polonica 64, 41-64 (doi: 10.4202/app.00569.2018). (Chambi-Trowell, S.A.V., Whiteside, D.I. and Benton, M.J.). pdf. Supplementary file.
A new species of Platysiagum from the Luoping Biota (Anisian, Middle Triassic, Yunnan, South China) reveals the relationship between Platysiagidae and Neopterygii. Geological Magazine 156, 669-682 (doi: 10.1017/S0016756818000079) (Wen, W., Hu, S.X., Zhang, Q.Y., Benton, M.J., Kriwet, J., Chen, Z.Q., Zhou, C.Y., Xie, T., and Huang, J.Y.). pdf. Supplementary file.
Gondolelloid multielement conodont apparatus (Nicoraella) from the middle Triassic of Yunnan Province, southwestern China. Palaeogeography, Palaeoclimatology, Palaeoecology 522, 98-110 (doi: 10.1016/j.palaeo.2018.07.015) (Huang, J.Y., Hu, S.X., Zhang, Q.Y., Donoghue, P.C.J., Benton, M.J., Zhou, C.Y., Martínez-Pérez, C., Wen, W., Xie, T., Chen, Z.-Q., Luo, M., Yao, H.Z., and Zhang, K.X.). pdf.
Morphological innovation and the evolution of hadrosaurid dinosaurs. Paleobiology 45, 347-362 (doi: 10.1017/pab.2019.9) (Stubbs, T.L., Benton, M.J., Elsler, A. and Prieto-Márquez, A.). pdf. Data available from the Dryad Digital Repository.
Ontogenetic stages of ceratopsian dinosaur Psittacosaurus in bone histology. Acta Palaeontologia Polonica 64, 323-334 (doi: 10.4202/app.00559.2018). (Zhao, Q., Benton, M.J., Hayashi, S. and Xu, X.). pdf. Supplementary file.
The stem group teleost Pachycormus (Pachycormiformes: Pachycormidae) from the Upper Lias (Lower Jurassic) of Strawberry Bank, UK. Paläontologische Zeitschrift 93, 285-302 (doi: 10.1007/s12542-018-0431-7) (Cawley, J.J., Kriwet, J., Klug, S., and Benton, M.J.). pdf.
Archibald Geikie and the Elgin reptiles. In Betterton, J., Craig, J., Mendum, J.R., Neller, R., and Tanner, J. (eds) Aspects of the Life and Works of Archibald Geikie. Geological Society, London, Special Publications 480, 353-359 (doi: 10.1144/SP480.4). pdf.
The Middle Triassic (Anisian) Otter Sandstone biota (Devon, UK): review, recent discoveries and ways ahead. Proceedings of the Geologists’ Association 130, 294-306 (doi: 10.1016/j.pgeola.2017.06.007) (Coram, R.A., Radley, J.D., and Benton, M.J.). pdf.
The Early Triassic Jurong fish fauna, South China: age, anatomy, taphonomy, and global correlation. Global & Planetary Change 180, 33-50 (doi: 10.1016/j.gloplacha.2019.05.012) (Qiu, X.C., Xu, Y.L., Chen, Z.-Q., Benton, M.J., Wen, W., and Huang, Y.C.). pdf.
Ontogenetic braincase development in Psittacosaurus lujiatunensis (Dinosauria: Ceratopsia) using micro-computed tomography. PeerJ 7, e7217 (doi: 10.7717/peerj.7217) (Bullar, C.M., Zhao, Q., Benton, M.J., and Ryan, M.J.). pdf.
The early origin of feathers. Trends in Ecology & Evolution 34, 856-869 (doi: 10.1016/j.tree.2019.04.018) (Benton, M.J., Dhouailly, D., Jiang, B.Y., and McNamara, M.). pdf.
Apparatus architecture of the conodont Nicoraella kockeli (Gondelelloidea, Prioniodinina) constrains functional interpretations. Palaeontology 62, 823-835 (doi: 10.1111/pala.12429) (Huang, J.Y., Martínez-Pérez, C., Hu, S.X., Zhang, Q.Y., Zhang, K.X., Zhou, C.Y., Wen, W., Xie, T., Benton, M.J., Chen, Z.-Q.,Luo, M., and Donoghue, P.C.J.). pdf. Data for this study are available here and here in the University of Bristol Digital Repository.
Diverse earliest Triassic ostracod fauna of the non-microbialite-bearing shallow marine carbonates of the Yangou section, South China. Lethaia 52, 583-596 (doi: 10.1111/let.12332) (Qiu, X., Tian, L., Wu, K., Benton, M.J., Sun, D., Yang, H. and Tong J.). pdf.
A new crurotarsan archosaur from the Late Triassic of South Wales. Journal of Vertebrate Paleontology 39, e1645147
(doi: 10.1080/02724634.2019.1645147) (Patrick, E.L., Whiteside, D.I. and Benton, M.J.). pdf.
Convergence and functional evolution of longirostry in crocodylomorphs. Palaeontology 62, 867-887 (doi: 10.1111/pala.12432) (Ballell, A., Moon, B.C., Porro, L.B., Benton, M.J., and Rayfield, E.J.). pdf. Data for this study are available in the Dryad Digital Repository.
Altered fluvial patterns in North China indicate rapid climate change linked to the Permian-Triassic mass extinction. Scientific Reports 9, 16818 (doi: 10.1038/s41598-019-53321-z) (Zhu, Z., Liu, Y.Q., Kuang, H.W., Benton, M.J., Newell, A.J., Xu, H., Ji, S.A., Xu, S.C., Peng, N., and Zhai, Q.G). pdf. Supplementary data.
Middle Triassic conodont apparatus architecture revealed by synchrotron X-ray microtomography. Palaeoworld 28, 429-440 (doi: 10.1016/j.palwor.2018.08.003) (Huang, J.Y., Martínez-Pérez, C., Hu, S.X., Donoghue, P.C.J., Zhang, Q.Y., Zhou, C.Y., Wen, W., Benton, M.J., Luo, M., Yao, H.Z., and Zhang, K.X.). pdf.
Microvertebrates from the Wadhurst Clay Formation (Lower Cretaceous) of Ashdown Brickworks, East Sussex, UK. Proceedings of the Geologists’ Association 130,752-769 (doi: 10.1016/j.pgeola.2019.08.003) (Turmine-Juhel, P., Wilks, R., Brockhurst, D., Austen, P.A., Duffin, C.J. and Benton, M.J.). pdf. Data for this study are available here.

2018

On formation-based sampling proxies and why they should not be used to correct the fossil record. Palaeontology 61, 119-132 (doi: 10.1111/pala.12331) (Dunhill, A.M., Hannisdal, B., Brocklehurst, N., and Benton, M.J.). pdf. Supplementary file, at Dryad.
First workshop on the Carnian Pluvial Episode (Late Triassic): a report. Albertiana 44, 49-57. (Dal Corso, J., Benton, M.J., Bernardi, M., Franz, M., Gianolla, P., Hohn, S., … Zhang, Y.) pdf.
Tetrapod distribution and temperature rise during the Permian-Triassic mass extinction. Proceedings of the Royal Society B 285, 20172331 (doi: 10.1098/rspb.2017.2331) (Bernardi, M., Petti, F.M., and Benton, M.J.). pdf. Supplement. Data.
Dinosaur diversification linked with the Carnian Pluvial Episode. Nature Communications 9, art. 1499 (doi: 10.1038/s41467-018-03996-1) (Bernardi, M., Gianolla, P., Petti, F. M., Mietto, P., and Benton, M.J.). pdf.
A new millipede (Diplopoda, Helminthomorpha) from the Middle Triassic Luoping biota of Yunnan, Southwest China. Journal of Paleontology 92, 478-487 (doi: 10.1017/jpa.2017.93) (Huang, J.Y., Hannibal, J.T., Feldmann, R.M., Zhang, Q.Y., Hu, S.X., Schweitzer, C.E., Benton, M.J., Zhou, C.Y., Wen, W., and Xie, T.).
The terrestrial fauna of the Late Triassic Pant-y-ffynnon Quarry fissures, South Wales, UK and a new species of Clevosaurus (Lepidosauria: Rhynchocephalia). Proceedings of the Geologists’ Association 129, 99-119 (doi: 10.1016/j.pgeola.2017.11.001) (Keeble, E., Whiteside, D.I., and Benton, M.J.). pdf. Supplementary file.
A Rhaetian microvertebrate fauna from Stowey Quarry, Somerset, U.K. Proceedings of the Geologists’ Association 129, 144-158 (doi: 10.1016/j.pgeola.2010.02.001) (Cavicchini, I., Heyworth, H.C., Duffin, C.J., Hildebrandt, C., and Benton, M.J.). pdf. Supplementary file.
Fossilized skin reveals coevolution with feathers and metabolism in feathered dinosaurs and early birds. Nature Communications 9, art. 2072 (doi: 10.1038/s41467-018-04443-x) (McNamara, M.E., Zhang, F., Kearns, S.L., Orr, P.J., Toulouse, A., Foley, T., Hone, D.W.E., Rogers, C.S., Benton, M.J., Johnson, D., Xu, X., and Zhou, Z.). pdf. Supplementary file.
Taxonomic reassessment of Clevosaurus latidens Fraser, 1993 (Lepidosauria, Rhynchocephalia) and rhynchocephalian phylogeny based on parsimony and Bayesian inference. Journal of Paleontology 92, 734-742 (doi: 10.1017/jpa.2017.136) (Herrera-Flores, J.A., Stubbs, T.L., Elsler, A., and Benton, M.J.). pdf.
Non-integumentary melanosomes can bias reconstructions of the colours of fossil vertebrates. Nature Communications 9, art. 2878 (doi: 10.1038//s41467-018-05148-x) (McNamara, M.E., Kaye, J.S., Benton, M.J., Orr, P.J., Rossi, V., Ito, S., and Wakamatsu, K.). pdf. Supplementary file.
Limuloid trackways from Permian-Triassic continental successions of North China. Palaeogeography, Palaeoclimatology, Palaeoecology 508, 71-90 (doi: 10.1016/j.palaeo.2018.07.022) (Shu, W.C., Tong, J.N., Benton, M.J., Chu, D.L., Yu, J.X., and Guo, W.W.). pdf.
Triassic extinctions and explosions. Geoscientist 28(8), 10-15 (Ruffell, A., Dal Corso, J., and Benton, M.J.). pdf.
Hyperthermal-driven mass extinctions: killing models during the Permian-Triassic mass extinction. Philosophical Transactions of the Royal Society, Series A 376, 20170076 (doi:10.1098/rsta.2017.0076). pdf.
Microvertebrates from the basal Rhaetian Bone Bed (latest Triassic) at Aust Cliff, S.W. England. Proceedings of the Geologists’ Association 129, 635-653 (doi: 10.1016/j.pgeola.2018.06.002) (Cross, S.R.R., Ivanovski, N., Duffin, C.J., Hildebrandt, C., Parker, A., and Benton, M.J.). pdf. Supplementary data.
Framboidal pyrite evidence for persistent low oxygen levels in shallow-marine facies of the Nanpanjiang Basin during the Permian-Triassic transition. Palaeogeography, Palaeoclimatology, Palaeoecology 511, 243-255 (doi: 10.1016/j.palaeo.2018.08.012) (Xiao, Y.F., Wu, K., Tian, L., Benton, M.J., Du, Y., Yang, H., and Tong, J.N.). pdf.
The Carnian Pluvial Episode and the origin of dinosaurs. Journal of the Geological Society 175, 1019-1026 (doi: 10.1144/jgs2018-049) (Benton, M.J., Bernardi, M., and Kinsella, C.). pdf. Supplementary file.
Multifaceted disparity approach reveals dinosaur herbivory flourished before the end-Cretaceous mass extinction. Paleobiology 44, 620-637 (doi: 10.1017/pab.2018.26) (Nordén, K.K., Stubbs, T.L., Prieto-Márquez, A. and Benton, M.J.). pdf.

2017

Live birth in an archosauromorph reptile. Nature Communications 8, art. 14445 (doi: 10.1038/ncomms14445) (Liu, J., Organ, C.L., Benton, M.J., Brandley, M.C. and Aitchison, J.C.). pdf. Supplementary file. Supplementary data. Article in ‘The Conversation’.
The impact of the Pull of the Recent on the fossil record of tetrapods. Evolutionary Ecology Research 18, 7-23 (Sahney, S. and Benton, M.J.). pdf. Supplementary file.
Response to: Phylogenetic placement, developmental trajectories and evolutionary implications of a feathered dinosaur tail in Mid-Cretaceous amber. Current Biology 27, R216-R217 (doi: 10.1016/j.cub.2016.10.008) (Xing, L., McKellar, R.C., Xu, X., Li, G., Bai, M., Persons, W.S., IV, Miyashita, T., Benton, M.J., Zhang, J.P., Wolfe, A.P., Yi, Q.R., Tseng, K.W., Ran, H., and Currie, P.J.). pdf.
Cellular preservation of musculoskeletal specializations in the Cretaceous bird Confuciusornis. Nature Communications 8, art. 14779 (doi: 10.1038/ncomms14779) (Jiang, B.Y., Zhao, T., Regnault, S., Edwards, N.P., Kohn, S.C., Li, Z.H., Wogelius, R.A., Benton, M.J. and Hutchinson, J.R.). pdf. Supplementary file.
Microbial mats in the terrestrial Lower Triassic of North China and implications for the Permian-Triassic mass extinction. Palaeogeography, Palaeoclimatology, Palaeoecology 474, 214-231 (doi: 10.1016/j.palaeo.2016.06.013) (Chu, D.L., Tong, J.N., Bottjer, D.J., Song, H.J., Song, H.Y., Benton, M.J., Tian, L., and Guo, W.W.). pdf
‘Residual diversity estimates’ do not correct for sampling bias in palaeodiversity data. Methods in Ecology and Evolution 8, 453-459 (doi: 10.1111/2041-210X.12666) (Sakamoto, M., Venditti, C., and Benton, M.J.). pdf. Supplementary file.
Taphonomy and palaeobiology of early Middle Triassic coprolites from the Luoping biota, southwest China: Implications for reconstruction of fossil food webs. Palaeogeography, Palaeoclimatology, Palaeoecology 474, 232-246 (doi: 10.1016/j.palaeo.2016.06.001) (Luo, M., Hu, S.X., Benton, M.J., Shi, G.R., Zhao, L.S., Huang, J.Y., Song, H.J., Wen, W., Zhang, Q.Y., Fang, Y.H., Huang, Y.G., and Chen, Z.Q.). pdf
Assessing sampling of the fossil record in a geographically and stratigraphically constrained dataset: the Chalk Group of Hampshire, southern UK. Journal of the Geological Society174, 509-521 (doi: 10.1144/jgs2016-093) (Walker, F.M., Dunhill, A.M., Woods, M.A., Newell, A.J., and Benton, M.J.). pdf. Supplementary file.
Macroevolutionary patterns in Rhynchocephalia: is the tuatara (Sphenodon punctatus) a living fossil? Palaeontology 60, 319-328 (doi: 10.1111/pala.12284) (Herrera-Flores, J., Stubbs, T.L., and Benton, M.J.). pdf. Supplementary file, at Dryad.
Biostratigraphy and geometric morphometrics of conchostracans (Crustacea, Branchiopoda) from the Late Triassic fissure deposits of Cromhall Quarry, UK. Palaeontology 60, 349-374 (doi: 10.1111/pala.12288) (Morton, J.D., Whiteside, D.I., Hethke, M., and Benton, M.J.). pdf. Supplementary file, at Dryad.
On the purported presence of fossilized collagen fibres in an ichthyosaur and a theropod dinosaur. Palaeontology 60, 409-422 (doi: 10.1111/pala.12292) (Smithwick, F.M., Mayr, G., Saitta, E.T., Benton, M.J., and Vinther, J.). pdf.
The first discovery of crinoids and cephalopod hooklets in the British Triassic. Proceedings of the Geologists’ Association 128, 360-373 (doi: 10.1016/j.pgeola.2017.03.005) (Landon, E.N.U., Duffin, C.J., Hildebrandt, C., Davies, T.G., Simms, M.J., and Benton, M.J.). pdf.
Leptolepid otoliths from the Hauterivian (Lower Cretaceous) Lower Weald Clay (southern England). Proceedings of the Geologists’ Association 128, 613-625 (doi: 10.1016/j.pgeola.2017.05.001) (Flannery Sutherland, J. T., Austen, P.A., Duffin, C.J., and Benton, M.J.). pdf.
Fossilization of soft tissues. National Science Review 4, 512-513. pdf.
Body length of bony fishes was not a selective factor during the biggest mass extinction of all time. Palaeontology 60, 727-741 (doi: 10.1111/pala.12309) (Puttick, M.N., Kriwet, J., Wen, W., Hu, S.X., Thomas, G.H., and Benton, M.J.). pdf. Supplementary file, at Dryad.
Exceptional appendage and soft-tissue preservation in a Middle Triassic horseshoe crab from SW China. Scientific Reports 7, 14112 (doi: 10.1038/s41598-017-13319-x) (Hu, S.-X., Zhang, Q.-Y., Feldmann, R.M., Benton, M.J., Schweitzer, C.E., Huang, J.Y., Wen, W., Zhou, C.Y., Xie, T., Lü, T. and Hong, S.G.). pdf. Supplementary file.
Untangling the dinosaur family tree. Nature 551, E1-E3 (doi: 10.1038/nature24011) (Langer, M.C., Ezcurra, M.D., Rauhut, O.W.M., Benton, M.J., Knoll, F., McPhee, B.W., Novas, F.E., Pol, D., and Brusatte, S.L.). pdf. Supplementary file.
Trophic and tectonic limits to the global increase of marine invertebrate diversity. Scientific Reports 7, 15969 (doi: 10.1038/s41598-017-16257-w) (Cermeño, P., Benton, M.J., Paz, O. and Vérard, C.). pdf.
Russia-UK collaboration in paleontology: past, present, and future. Paleontological Journal 51, 576-599 (doi: 10.1134/S0031030117060028) (Benton, M.J., Briggs, D.E.G., Clack, J.A., Edwards, D., Galway-Witham, J., Stringer, C.B., and Turvey, S.). pdf.

2016

Digit-only sauropod pes trackways from China – evidence of swimming or a preservational phenomenon? Scientific Reports 6, article no. 21138 (doi: 10.1038/srep21138) (Xing, L.D., Li, D.Q., Falkingham, P.L., Lockley, M.G., Benton, M.J., Klein, H., Zhang, J.P., Ran, H., Persons, W.S., and Dai, H.). pdf. Supplementary file.
The challenges to inferring the regulators of biodiversity in deep time. Philosophical Transactions of the Royal Society, Series B 371, 20150216 (doi: 10.1098/rstb.2015.0216) (Ezard, T.H.G., Quental, T.B., and Benton, M.J.) pdf.
Radiation and extinction: investigating clade dynamics in deep time. Biological Journal of the Linnean Society 118, 6-12 (doi: 10.1111/bij.12763) (Goswami, A., Mannion, P.D., and Benton, M.J.). pdf.
The Rhaetian vertebrates of Chipping Sodbury, South Gloucestershire, UK, a comparative study. Proceedings of the Geologists’ Association 127, 40-52 (doi: 10.1016/j.pgeola.2016.02.010) (Lakin, R.J., Duffin, C.J., Hildebrandt, C., and Benton, M.J.). pdf. Supplementary file.
Dating Placentalia: morphological clocks fail to close the molecular-fossil gap. Evolution 70, 873-886 (doi: 10.1111/evo.12907) (Puttick, M.N., Thomas, G.H., and Benton, M.J.). pdf. Supplementary file.
Dinosaurs in decline tens of millions of years before their final extinction. Proceedings of the National Academy of Sciences, U.S.A. 113, 5036-5040 (doi: 10.1073/pnas.1521478113) (Sakamoto, M., Benton, M.J., and Venditti, C.). pdf. Article in ‘The Conversation’.
A new Minisauripus site from the Lower Cretaceous of China: Tracks of small adults or juveniles? Palaeogeography, Palaeoclimatology, Palaeoecology 452, 28-39 (doi: 10.1016/j.palaeo.2016.04.006) (Xing, L., Lockley, M.G., Yang, G., Cao, J., Benton, M.J., Xu, X., Zhang, J.P., Klein, H., Persons, W.S., IV, Kim, J.Y., Peng, G.Z., Ye, Y., and Ran, H.). pdf.
The Late Triassic and Early Jurassic fissure faunas from Bristol and South Wales. Palaeontologia Polonica 67, 257-287 (Whiteside, D.I., Duffin, C.J., Gill, P.G., Marshall, J.E.A., and Benton, M.J.) (doi: 10.4202/pp.2016.67_257). pdf.
Mummified precocial bird wings in mid-Cretaceous Burmese amber. Nature Communications 7, 12089 (doi: 10.1038/ncomms12089) (Xing, L., McKellar, R.C., Wang, M., Bai, M., O’Connor, J.K., Benton, M.J., Zhang, J.P., Wang, Y., Tseng, K.W., Lockley, M.G., Li, G., Zhang, W.W., and Xu, X.). pdf. Supplementary data. Nature news story. Article in ‘The Conversation’.
Dynamics of dental evolution in ornithopod dinosaurs. Scientific Reports 6, 28904 (doi: 10.1038/srep28904) (Strickson, E., Prieto-Márquez, A., Benton, M.J., and Stubbs, T.L.). pdf. Supplementary data 1. Supplementary data 2.
The Chinese pareiasaurs. Zoological Journal of the Linnean Society 127, 813-853 (doi: 10.1111/zoj.12389). pdf.
Severe selenium depletion in the Phanerozoic oceans as a factor in three global mass extinction events. Gondwana Research 36, 209-218 (doi: 10.1016/j.gr.2015.10.001) (Long, J.A., Large, R.R., Lee, M.S.Y., Benton, M.J., Danyushevsky, L.V., Chiappe, L.M., Halpin, J.A., Cantrill, D., and Lottermoser, B.). pdf. Read a discussion at ‘The Conversation’ by John Long here.
Microvertebrates from multiple bone beds in the Rhaetian of the M4-M5 motorway junction, South Gloucestershire, U.K. Proceedings of the Geologists’ Association 127, 464-477 (doi: 10.1016/j.pgeola.2016.07.001) (Slater, T.S., Duffin, C.J., Hildebrandt, C., Davies, T.G., and Benton, M.J.). pdf. Supplementary file.
The Rhaetian (Late Triassic) vertebrates of Hampstead Farm Quarry, Gloucestershire, UK. Proceedings of the Geologists’ Association 127, 478-505 (doi: 10.1016/j.pgeola.2016.05.003) (Mears, E.M., Rossi, V., MacDonald, E., Coleman, G., Davies, T.G., Arias-Riesgo, C., Hildebrandt, C., Thiel, H., Duffin, C.J., Whiteside, D.I., and Benton, M.J.). pdf. Supplementary file.
Belowground rhizomes in paleosols: The hidden half of an Early Devonian vascular plant. Proceedings of the National Academy of Sciences, U.S.A. 113, 9451-9456 (doi: 10.1073/pnas.1605051113) (Xue, J.Z., Deng, Z.Z., Huang, P., Huang, K.J., Benton, M.J., Cui, Y., Wang, D.M., Liu, J.B., Shen, B., Basinger, J.F., and Hao, S.G.). pdf. Supplementary file.
Fish and tetrapod communities across a marine to brackish salinity gradient in the Pennsylvanian (early Moscovian) Minto Formation of New Brunswick, Canada, and their palaeoecological and palaeogeographical implications. Palaeontology 59, 689-724 (doi: 10.1111/pala.12249) (Ó Gogáin, A., Falcon-Lang, H.J., Carpenter, D.K., Miller, R.F., Benton, M.J., Pufahl, P.K., Ruta, M., Davies, T.G., Hinds, S.J., and Stimson, M.R.). pdf.
Biostratigraphic correlation and mass extinction during the Permian-Triassic transition in terrestrial-marine siliciclastic settings of South China. Global & Planetary Change 146, 67-88 (doi: 10.1016/j.gloplacha.2016.09.009) (Chu, D.L., Yu, J.X., Tong, J.N., Benton, M.J., Song, H.J., Huang, Y.F., Song, T., and Tian, L.). pdf
Ecomorphological diversifications of Mesozoic marine reptiles: the roles of ecological opportunity and extinction. Paleobiology 42, 547-573 (doi: 10.1017/pab.2016.15) (Stubbs, T.L. and Benton, M.J.). pdf.
Palaeontology: dinosaurs, boneheads and recovery from extinction. Current Biology 26, R882-R902 (doi: 10.1016/j.cub.2016.07.029). pdf.
Origins of biodiversity. PLoS Biology 14(11), e2000724 (doi: 10.1371/journal. pbio.2000724). pdf. In the top-50 most downloaded PLoS Biology articles of 2016.
The Triassic. Current Biology 26, R1214-R1218 (doi: 10.1016/j.cub.2016.10.060). pdf.
A feathered dinosaur tail with primitive plumage trapped in mid-Cretaceous amber. Current Biology 26, 3352-3360 (doi: 10.1016/j.cub.2016.10.008) (Xing, L., McKellar, R.C., Xu, X., Li, G., Bai, M., Persons, W.S., IV, Miyashita, T., Benton, M.J., Zhang, J.P., Wolfe, A.P., Yi, Q.R., Tseng, K.W., Ran, H., and Currie, P.J.). pdf. Supplementary file. Selected as the Number 1 scientifically influential fossil story of 2016 by Earth Archives. Top-ten Altmetric score for 2017, and highest-ever Altmetric score for Current Biology.

2015

Latest Triassic marine sharks and bony fishes from a bone bed preserved in a burrow system, from Devon, UK. Proceedings of the Geologists’ Association 126, 130-142 (doi: 10.1016/j.pgeola.2014.11.004) (Korneisel, D., Gallois, R.W., Duffin, C.J., and Benton, M.J.). pdf.
Constraints on the timescale of animal evolutionary history. Palaeontologia Electronica 18.1.1FC, 1-106 (Benton, M.J., Donoghue, P.C.J., Asher, R.J., Friedman, M., Near, T.J., and Vinther, J.). pdf.
Mesozoic echinoid diversity in Portugal: investigating fossil record quality and environmental constraints on a regional scale. Palaeogeography, Palaeoclimatology, Palaeoecology424, 132-146 (doi: 10.1016/j.palaeo.2015.02.014) (Pereira, B.C., Benton, M.J., Ruta, M., and Mateus, O.). pdf.
The Chinese Pompeii? Death and destruction of dinosaurs in the Early Cretaceous of Lujiatun, NE China. Palaeogeography, Palaeoclimatology, Palaeoecology 427, 89-99 (doi: 10.1016/j.palaeo.2015.03.037) (Rogers, C.S., Hone, D.W.E., McNamara, M.E., Zhao, Q., Orr, P.J., Kearns, S.L., and Benton, M.J.). pdf.
The fossil record of ichthyosaurs, completeness metrics and sampling biases. Palaeontology 58, 521-536 (doi: 10.1111/pala.12158) (Cleary, T.J., Moon, B.C., Dunhill, A.M., and Benton, M.J.) pdf. Supplementary file. All data and supplements are located here.
A re-evaluation of goniopholidid crocodylomorph material from Central Asia: biogeographic and phylogenetic implications. Acta Palaeontologica Polonica 60, 291-312 (doi: 10.4202/app.2013.0018) (Halliday, T.J.D., Andrade, M.B. de, Benton, M.J., and Efimov, M.B.). pdf. Supplementary file
Early Triassic wrinkle structures on land: stressed environments and oases for life. Scientific Reports 5, 10109 (doi: 10.1038/srep10109) (Chu, D.L., Tong, J.N., Song, H.J., Benton, M.J., Bottjer, D.J., Song, H.Y., and Tian, L.). pdf.
Stepwise evolution of Palaeozoic tracheophytes from South China: Contrasting leaf disparity and taxic diversity. Earth-Science Reviews 148, 77-93 (doi: 10.1016/j.earscirev.2015.05.013) (Xue, J.Z., Huang, P., Ruta, M., Benton, M.J., Hao, S.G., Xiong, C.H., Wang D.M., Cascales-Miñana, B., Wang, Q., and Liu, L.). pdf.
Exploring macroevolution using modern and fossil data. Proceedings of the Royal Society B 282, 20150569. (doi: 10.1098/rspb.2015.0569) (Benton, M.J.). pdf.
A distinctive Late Triassic microvertebrate fissure fauna and a new species of Clevosaurus (Lepidosauria: Rhynchocephalia) from Woodleaze Quarry, Gloucestershire, UK. Proceedings of the Geologists’ Association 126, 402-416 (doi: 10.1016/j.pgeola.2015.05.003) (Klein, C.G., Whiteside, D.I., Selles de Lucas, V., Viegas, P.A., and Benton, M.J.) pdf.
Lilliput effect in freshwater ostracods during the Permian-Triassic extinction. Palaeogeography, Palaeoclimatology, Palaeoecology 435, 38-52 (doi: 10.1016/j.palaeo.2015.06.003) (Chu, D.L., Tong, J.N., Song, H.J., Benton, M.J., Song, H.Y., Qiu, X.C., Huang, Y.F., and Tian, L.). pdf. Supplementary file
Early Pennsylvanian (Langsettian) fish assemblages from the Joggins Formation, Canada, and their implications for palaeoecology and palaeogeography. Palaeontology 58, 661-690 (doi: 10.1111/pala.12164) (Carpenter, D.K., Falcon-Lang, H.J., Benton, M.J., and Grey, M.). pdf.
The skull and endocranium of a Lower Jurassic ichthyosaur based on digital reconstructions. Palaeontology 58, 732-745 (doi: 10.111/pala.12174) (Marek, R.D., Moon, B.C., Williams, M., and Benton, M.J.). pdf. Supplementary file
The Fossil Calibration Database, a new resource for divergence dating. Systematic Biology 65, 853-859 (doi: 10.1093/sysbio/syv025)(Ksepka, D.T., Parham, J.F., Allman, J.F., Benton, M.J., Carrano, M.T., Cranston, K.A., Donoghue, P.C.J., Head, J.J., Hermsen, E.J., Irmis, R.B., Joyce, W.G., Kohli, M., Lamm, K.S., Leehr, D., Patané, J.S.L., Polly, P.D., Phillips, M.J., Smith, N.A., Smith, N.D., Van Tuinen, M., Ware, J.L., and Warnock, R.C.M.). pdf.
Complete biotic and sedimentary records of the Permian-Triassic transition from Meishan section, South China: Ecologically assessing mass extinction and its aftermath. Earth-Science Reviews, 149, 67-107 (doi: 10.1016/j.earscirev.2014.10.005) (Chen, Z.-Q., Yang, H., Luo, M., Benton, M.J., Kaiho, K., Zhao, L.S., Huang, Y.G., Zhang, K.X., Fang, Y.H., Jiang, H.S., Qiu, H., Li, Y., Tu, C.Y., Shi, L., Zhang, L., Feng, Z.Q., and Chen, L.). pdf.
A marine vertebrate fauna from the Late Triassic of Somerset, and a review of British placodonts. Proceedings of the Geologists’ Association 126, 564-581 (doi: 10.1016/j.pgeola.2015.07.001) (Nordén, K.K., Duffin, C.J., and Benton, M.J.). pdf.
Palaeodiversity and formation counts: redundancy or bias? Palaeontology 58, 1003-1029 (doi: 10.1111/pala.12191). pdf. Download the data here.
The Strawberry Bank Lagerstätte reveals insights into Early Jurassic life. Journal of the Geological Society 172, 683-692 (doi: 10.1144/jgs2014-144) (Williams, M., Benton, M.J., and Ross, A.). pdf.
Reply to the comment on Chu et al., “Lilliput effect in freshwater ostracods during the Permian-Triassic extinction” [Palaeogeography, Palaeoclimatology, Palaeoecology 435 (2015): 38-52. Palaeogeography, Palaeoclimatology, Palaeoecology 440, 863-865 (doi: : 10.1016/j.palaeo.2015.09.018) (Chu, D.L., Tong, J.N., Song, H.J., Benton, M.J., Song, H.Y., Qiu, X.C., Huang, Y.F., and Tian, L.). pdf.
Microvertebrates from the classic Rhaetian bone beds of Manor Farm Quarry, near Aust (Bristol, UK). Proceedings of the Geologists’ Association 126, 762-776 (doi: 10.1016/j.pgeola.2015.09.002) (Allard, H., Carpenter, S.C., Duffin, C.J., and Benton, M.J.). pdf. Supplementary file

2014

Marine flooding event in continental Triassic facies identified by a nothosaur and placodont bonebed (South Iberian Paleomargin). Facies 60, 277-293 (doi: 10.1007/s10347-013-0360-6) (Reolid, M., Pérez-Valera, F., Benton, M.J.). pdf.
Models for the rise of the dinosaurs. Current Biology 24, R87-R95 (doi: 10.1016/j.cub.2013.11.063) (Benton, M.J., Forth, J., and Langer, M.C.). pdf.
Impacts of global warming on Permo-Triassic terrestrial ecosystems. Gondwana Research 25, 1308-1337 (doi: 10.1016/j.gr.2012.12.010) (Benton, M.J. and Newell, A.J.) pdf.
Testing the fossil record: sampling proxies and scaling in the British Triassic-Jurassic. Palaeogeography, Palaeoclimatology, Palaeoecology 404, 1-11 (doi: 10.1016/j.palaeo.2014.03.026) (Dunhill, A.M., Benton, M.J., Twitchett, R.J., and Newell, A.J.) pdf.
High rates of evolution preceded the origin of birds. Evolution 68, 1497-1510 (doi: 10.1111/evo.12363) (Puttick, M.N., Thomas, G.H., and Benton, M.J.). pdf. Supplementary file. Working data.
Complex rostral neurovascular system in a giant pliosaur. Naturwissenschaften 101, 453-456 (doi: 10.1007/s00114-014-1173-3) (Foffa, D., Sassoon, J., Cuff, A.R., Mavrogordato, M.N., and Benton, M.J.) pdf. This was discussed in a story in New Scientist here.
Nothosaur foraging tracks from the Middle Triassic of southwestern China. Nature Communications 5, 3973 (doi:10.1038/ncomms4973) (Zhang, Q.Y., Wen, W., Hu, S.X., Benton, M.J., Zhou, C.Y., Xie, T., Lü, T., Huang, J.Y., Choo, B., Chen, Z.-Q., Liu, J., and Zhang, Q.C.). pdf. Supplement pdf.
Vertebrates from the Late Triassic Thecodontosaurus-bearing rocks of Durdham Down, Clifton (Bristol, UK). Proceedings of the Geologists’ Association 125, 317-328 (doi: 10.1016/j.pgeola.2014.02.002) (Foffa, D., Whiteside, D.I., Viegas, P. A., and Benton, M.J.) pdf.
Functional anatomy and feeding biomechanics of a giant Upper Jurassic pliosaur (Reptilia: Sauropterygia) from Weymouth Bay, Dorset, UK. Journal of Anatomy 225, 209-219 (doi: 10.1111/joa.12200) (Foffa, D., Cuff, A.R., Sassoon, J., Rayfield, E.J., Mavrogordato, M.N., and Benton, M.J.) pdf. Supplementary file.
A Jurassic ornithischian dinosaur from Siberia with both feathers and scales. Science 345, 451-455 (doi: 10.1126/science.1253351) (Godefroit, P., Sinitsa, S.M., Dhouailly, D., Bolotsky, Y.L., Sizov, A.V., McNamara, M.E., Benton, M.J., and Spagna, P.). pdf. Supplementary file.
Evolution: How birds became birds. Science 345, 508-509 (doi: 10.1126/science.1257633). pdf.
Sea surface temperature contributes to marine crocodylomorph evolution. Nature Communications 5, 4658 (doi: 10.1038/ncomms5658) (Martin, J.,E., Amiot, C., Lécuyer, and Benton, M.J.). pdf. Download Supplementary Excel file.
Disentangling rock record bias and common-cause from redundancy in the British fossil record. Nature Communications 5, 4818 (doi: 10.1038/ncomms5818) (Dunhill, A.M., Hannisdal, B.J., and Benton, M.J.). pdf. Download Supplementary file. Download Supplementary Excel file.
Filling the ceratosaur gap: A new ceratosaurian theropod from the Early Cretaceous of Spain. Acta Palaeontologica Polonica 59, 581-600 (doi: 10.4202/app.2011.0144) (Sánchez-Hernández, B. and Benton, M.J.). pdf.
The Bristol Dinosaur Project – a conservation and preparation overview. Journal of Paleontological Techniques 13, 50-64 (Viegas, P.A. and Benton, M.J.). pdf.
Response to Comment on “A Jurassic ornithischian dinosaur from Siberia with both feathers and scales”. Science 346, 343-c (doi: 10.1126/science.1260146) (Godefroit, P., Sinitsa, S.M., Dhouailly, D., Bolotsky, Y.L., Sizov, A.V., McNamara, M.E., Benton, M.J., and Spagna, P.). pdf.
Carboniferous (Tournaisian) fish assemblages from the Isle of Bute, Scotland: systematics and palaeoecology. Palaeontology 57, 1215-1240 (doi: 10.1111/pala.12112) (Carpenter, D.K., Falcon-Lang, H.J., Benton, M.J., and Henderson, E.). pdf.
A gigantic nothosaur (Reptilia: Sauropterygia) from the Middle Triassic of SW China and its implication for the Triassic biotic recovery. Nature Scientific Reports 4, 7142 (doi: 10.1038/srep07142) (Liu, J., Hu, S.X., Rieppel, O., Jiang, D.Y., Benton, M.J., Kelley, N.P., Aitchison, J.C., Zhou,C.Y., Wen, W., Huang, J.Y., Xie, Y., and Lv, T.). pdf.
Juvenile-only clusters and behaviour of the Early Cretaceous dinosaur Psittacosaurus. Acta Palaeontologica Polonica 59, 827-833 (doi: 10.4202/app.2012.0128) (Zhao, Q., Benton, M.J., Xu, X., and Sander, M.J.). pdf. Supplementary file
Defining the discipline of geobiology. National Science Review 1, 483-485 (doi: 10.1093/nsr/nwu052) (Benton, M.J. and Xie, S.C.). pdf.

2013

Origins of biodiversity. Palaeontology 56, 1-7 (doi: 10.1111/pala.12012). pdf.
The first half of tetrapod evolution, sampling proxies, and fossil record quality. Palaeogeography, Palaeoclimatology, Palaeoecology 372, 18-41 (doi: 10.1016/j.palaeo.2012.09.005) (Benton, M.J., Ruta, M., Dunhill, A.M., and Sakamoto, M.). pdf. The Early Tetrapod Database. Supporting data.
Coelacanths from the Middle Triassic Luoping Biota, Yunnan, South China, with the earliest evidence of ovoviviparity. Acta Palaeontologica Polonica 58, 175-193 (doi: 10.4202/app.2011.0066) (Wen, W., Zhang, Q.Y., Hu, S.X., Benton, M.J., Zhou, C.Y., Xie, T., Huang, J.-Y., and Chen, Z.Q.). pdf.
Completeness of the fossil record and the validity of sampling proxies: a case study from the Triassic of England and Wales. Journal of the Geological Society 170, 291-300 (Dunhill, A.M., Benton, M.J., Newell, A.J., and Twitchett, R.J.) (doi: 10.1144/jgs2012-025). pdf. Supporting data (compressed folder).
Primary feather lengths may not be important for inferring the flight styles of Mesozoic birds. Lethaia 46, 146-153 (doi: 10.1111/j.1502-3931.2012.00325.x) (Chan, N.J., Dyke, G.J., and Benton, M.J.). pdf. Supporting data.
Histology and postural change during the growth of the ceratopsian dinosaur Psittacosaurus lujiatunensis. Nature Communications 4, 2079 (doi: 10.1038/ncomms3079) (Zhao, Q., Benton, M.J., Sullivan, C., Sander, P.M., and Xu, X.). pdf. Supporting data.
The first specimen of the Middle Triassic Phalarodon atavus (Ichthyosauria: Mixosauridae) from South China, showing postcranial anatomy and peri-Tethyan distribution. Palaeontology 56, 849-866 (doi: 10.1111/pala.12021) (Liu, J., Motani, R., Jiang, D.Y., Hu, S.X., Aitchison, J.C., Rieppel, O., Benton, M.J., Zhang, Q.Y., Zhou, C.Y.). pdf.
A new shrimp (Decapoda, Dendrobranchiata, Penaeoidea) from the Middle Triassic of Yunnan, Southwest China. Journal of Paleontology 87, 603-611 (doi: 10.1666/13-024). (Huang, J., Feldmann, R.M., Schweitzer, C.E., Hu, S.H., Zhou, C.Y., Benton, M.J., Zhang, Q.Y., Wen, W., Xie, T.). pdf.
Carbonate reticulated ridge structures from the Lower Middle Triassic of the Luoping area, Yunnan, southwestern China: geobiologic features and implications for exceptional preservation of the Luoping biota. Palaios 28, 541-551 (doi: 10.2110/palo.2012.p12-122r) (Luo, M., Chen, Z.Q., Hu, S., Zhang, Q.Y., Benton, M.J., Zhou, C.Y., Wen, W., Huang, J.Y.) pdf.
No gap in the Middle Permian record of terrestrial vertebrates: REPLY. Geology 41, e294 (doi: 10.1130/G34595Y.1). pdf.
Decoupling of morphological disparity and taxic diversity during the adaptive radiation of anomodont therapsids. Proceedings of the Royal Society B 280, 20131071 (doi: 10.1098/rspb.2013.1071) (Ruta, M., Angielczyk, K.D., Fröbisch, J., Benton, M.J.). pdf. Supplementary data explanation. Supplementary data; Excel file.
The radiation of cynodonts and the ground plan of mammalian morphological diversity. Proceedings of the Royal Society B 280: 20131865. (Ruta, M., Botha-Brink, J., Mitchell, S.A., Benton, M.J.). (doi: 10.1098/rspb.2013.1865). pdf. Supplementary data explanation. Supplementary data; Excel file. Discussion by us at The Conversation.
Exceptional vertebrate biotas from the Triassic of China, and the expansion of marine ecosystems after the Permo-Triassic mass extinction. Earth-Science Reviews 123, 199-243 (doi: 10.1016/j.earscirev.2013.05.014) (Benton, M.J., Zhang, Q.Y., Hu, S.X,, Chen, Z.Q., Wen, W., Liu, J., Huang, J.Y., Zhou, C.Y., Xie, T., Tong, J.N., and Choo, B.) pdf. For further images and commentary of Brian Choo’s spectacular paintings for this paper, read more about the Luoping, Xingyi, and Guanling images.
Diversity dynamics of Silurian-Early Carboniferous land plants in South China. PLoS ONE 8(9): e75706. (doi:10.1371/journal.pone.0075706) (Xiong, C.H., Wang, D.M., Wang, Q., Benton, M.J., Xue, J.H., Meng, M.C., Zhao, Q., Zhang, J.). pdf.
Causes and consequences of mass extinction. In Losos, J.B. (ed.) The Princeton Guide to Evolution, Princeton University Press, Princeton and Oxford, pp. 580-585. pdf. Explore the remainder of the book, or buy a copy here.

2012

The Bristol Dinosaur Project. Proceedings of the Geologists’ Association 123, 210-225 (doi:10.1016/j.pgeola.2011.07.004) (Benton, M.J., Schouten, R., Drewitt, E.J.A., and Viegas, P.). pdf. View the official Bristol Dinosaur Project web site.
Preservation of exceptional vertebrate assemblages in Middle Permian fluviolacustrine mudstones of Kotel’nich, Russia: stratigraphy, sedimentology, and taphonomy. Palaeogeography, Palaeoclimatology, Palaeoecology 319-320, 58-83 (doi: 10.1016/j.palaeo.2012.01.005) (Benton, M.J., Newell, A.J., Khlyupin, A.Yu., Shumov, I.S., Price, G.D., and Kurkin, A.A.). pdf. Read our web pages about Kotel’nich here.
Best practices for justifying fossil calibrations. Systematic Biology 61, 346-359 (doi: 10.1093/sysbio/syr107) (Parham, J.F., Donoghue, P.C.J., Bell, C.J., Calway, T.D., Head, J.J., Holroyd, P.A., Inoue, J.G., Irmis, R.B., Joyce, W.G., Ksepka, D.T., Patané, J.S.L., Smith, N.D., Tarver, J.E., Van Tuinen, M., Yang, Z., Angielczyk, K.D., Greenwood, J.M., Hipsley, C.A., Jacobs, L., Makovicky, P.J., Müller, J., Smith, K.T., Theodor, J.M., Warnock, R.C.M., and Benton, M.J.) pdf.
A new genus of actinopterygian fish from the Anisian (Middle Triassic) of Luoping, Yunnan Province, Southwest China. Acta Palaeontologica Polonica 57, 149-160 (doi: 10.4202/app.2010.0089) (Wen, W., Zhang, Q.Y., Hu, S.X., Zhou, C.Y., Huang, J.Y., Chen, Z.Q., and Benton, M.J.) pdf.
Obituary: Arthur Richard Ivor Cruickshank 1932-2011; palaeontologist who specialized in the dicynodonts of Gondwana and Jurassic plesiosaurs. Geoscientist 22 (2), 28 (Taylor, M.A. and Benton, M.J). pdf.
Obituary: Arthur Cruickshank – 1932-2011. Palaeontologica Africana 46, 93-98 (Taylor, M.A., Benton, M.J., Noè, L.F., and Fraser, N.C.). pdf.
No gap in the Middle Permian record of fossil vertebrates. Geology 40, 339-342 (doi: 10.1130/G32669.1). pdf. Supplementary data.
The timing and pattern of biotic recovery following the end-Permian mass extinction. Nature Geoscience 5, 375-383 (doi:10.1038/ngeo1475) (Chen, Z.Q. and Benton, M.J.) pdf.
Origin and evolution of dinosaurs. 336-351 In Brett-Surman, M.K., Holtz, T.R., Jr., Farlow, J.O., and Walters, B. (eds), The complete dinosaur, 2nd edition. Indiana University Press, Bloomington, 1115 pp. pdf.
Cranial anatomy, taxonomic implications and palaeopathology of an upper Jurassic pliosaur (Reptilia: Sauropterygia) from Westbury, Wiltshire UK. Palaeontology 55, 743-773 (doi: 10.1111/j.1475-4983.2012.01151.xx) (Sassoon, J., Noè, L.F., and Benton, M.J.). pdf.
Calcretes, fluviolacustrine sediments and subsidence patterns in Permo-Triassic salt-walled minibasins of the south Urals, Russia. Sedimentology 59, 1659-1676 (doi: 10.1111/j.1365-3091.2012.01320.x) (Newell, A.J., Benton, M.J., Kearsey, T., Taylor, G.K., Twitchett, R.J., and Tverdokhlebov, V.P.). pdf.
The Late Triassic microvertebrate fauna of Tytherington, UK. Proceedings of the Geologists’ Association 123, 638-648 (doi: 10.1016/j.pgeola.2012.05.003) (Van den Berg, T., Whiteside, D.I., Viegas, P., Schouten, R., and Benton, M.J.). pdf. Supplementary information (specimen list).
Naming the Bristol dinosaur, Thecodontosaurus: politics and science in the 1830s. Proceedings of the Geologists’ Association 123, 766-778 (doi: 10.1016/j.pgeola.2012.07.012). pdf. Brian Switek’s article about this paper, for the Smithsonian Museum, Tussling over Thecodontosaurus
Grit not grass: concordant patterns of early origin of hypsodonty in Great Plains ungulates and Glires. Palaeogeography, Palaeoclimatology, Palaeoecology 365-366, 1-10 (doi: 10.1016/j.palaeo.2012.09.001) (Jardine, P.E., Janis, C.M., Sahney, S., and Benton, M.J.). pdf.
Completeness of the fossil record and the validity of sampling proxies at outcrop level. Palaeontology 55, 1155-1175 (doi: 10.1111/j.1475-4983.2012.01149.x). (Dunhill, A.M., Benton, M.J., Twitchett, R.J., and Newell, A.J.). pdf. Supporting data.

2011

Archosaur remains from the Otter Sandstone Formation (Middle Triassic, late Anisian) of Devon, southern UK. Proceedings of the Geologists’ Association 122, 25-33 (doi: 10.1016/j.pgeola.2010.08.004). pdf.
Early Cretaceous (Berriasian) birds and pterosaurs from the Cornet bauxite mine, Romania. Palaeontology 54, 79-95 (doi: 19.1111/j.1475-4983.2010.00997.x) (Dyke, G.J., Benton, M.J., Posmosanu, E., and Naish, D.). pdf. Darren Naish’s blog on the birds and pterosaurs give a popular commentary.
Instant Expert 9: Mass extinctions. New Scientist 209 (2802), i-viii. pdf.
Is evolutionary history repeatedly rewritten in light of new fossil discoveries? Proceedings of the Royal Society, Series B 278, 599-604 (doi: 10.1098/rspb.2010.0663) (Tarver, J.E., Donoghue, P.C.J., and Benton, M.J.) pdf. Read more here, and on the Nature and Discover websites.
Crocodylomorph eggs and eggshells from the Adamantina Formation (Bauru Group), Upper Cretaceous of Brazil. Palaeontology 54, 309-321 (doi:10.1111/j.1475-4983.2010.01028.x). (Oliveira, C.E.M., Santucci, R.M., Andrade, M.B., Fulfaro, V.J., Basílio, J.A.F., and Benton, M.J). pdf.
Saltopus, a dinosauriform from the Upper Triassic of Scotland. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 101, 285-299 (doi: 10.1017/S1755691011020081) (Benton, M.J. and Walker, A.D.). pdf.
Macroevolutionary patterns in the evolutionary radiation of archosaurs (Tetrapoda: Diapsida). Earth and Environmental Science Transactions of the Royal Society of Edinburgh 101, 367-382 (doi: 10.1017/S1755691011020056) (Brusatte, S.L., Benton, M.J., Lloyd, G.T., Ruta, M., and Wang, S.C.). pdf.
Resetting the evolution of marine reptiles at the Triassic-Jurassic boundary. Proceedings of the National Academy of Sciences of the USA 108, 8339-8344 (doi: 10.1073/pnas.1018959108) (Thorne, P.M., Ruta, M., and Benton, M.J.). pdf of paper, and pdf of supplementary materials. View explanatory web site.
The Luoping biota: exceptional preservation, and new evidence on the Triassic recovery from end-Permian mass extinction. Proceedings of the Royal Society, Series B 278, 2274-2282 (doi: 10.1098/rspb.2010.2235) (Hu, S.-x., Zhang, Q.-y., Chen, Z.-Q., Zhou, C.y., Lü, T., Xie, T, Wen, W., Huang, J.-y., and Benton, M.J.). pdf.
Evolution of morphological disparity in pterosaurs. Journal of Systematic Palaeontology 9, 337-353 (doi: 10.1080/14772019.2011.565081). (Prentice, K.C., Ruta, M. and Benton, M.J.). pdf. View explanatory web site.
Ichthyosauria from the Upper Lias of Strawberry Bank, England. Palaeontology 54, 1069-1093. (doi: 10.1111/j.1475-4983.2011.01093.x). (Caine, H. and Benton, M.J.). pdf.
Fishes and tetrapods in the Upper Pennsylvanian (Kasimovian) Cohn Coal Member of the Mattoon Formation of Illinois, United States: systematics, paleoecology, and paleoenvironments. Palaios 26, 639-657 (doi: 10.2110/palo.2010.p10-121r). (Carpenter, D., Falcon-Lang, H.J., Benton, M.J., and Nelson, W.J.) pdf.
A new Late Jurassic turtle from Spain: phylogenetic implications, taphonomy and palaeoecology. Palaeontology 54, 1393-1414 (doi: 10.1111/j.1475-4983.2011.01100.x) (Slater, B.J., Reolid, M., Schouten, R., and Benton, M.J.). pdf.
Synthesizing and databasing fossil calibrations: divergence dating and beyond. Biology Letters 7, 801-803 (doi: 10.1098/rsbl.2011.0356) (Ksepka, D.T., Benton, M.J., Carrano, M.T., Gandolfo, M.A., Head, J.J., Hermesen, E.J., Joyce, W.G., Lamm, K.S., Patané, J.S.L., Phillips, M.J., Polly, P.D., Van Tuinen, M., Ware, J.L., Warnock, R.C.M., and Parham, J.F.). pdf.
Assessing the quality of the fossil record: insights from vertebrates. In: McGowan, A.J. & Smith, A.B. (eds) Comparing the Geological and Fossil Records: Implications for Biodiversity Studies. Geological Society, London, Special Publications 358, 63-94 (doi: 10.1144/SP358.6) (Benton, M.J., Dunhill, A.M., Lloyd, G.T. and Marx, F.G.). pdf.
A new Berriasian species of Goniopholis (Mesoeucrocodylia, Neosuchia) from England,and a review of the genus. Zoological Journal of the Linnean Society 163, S66-S108 (doi: 10.1111/j.1096-3642.2011.00709.x) (Andrade, M.B. de, Edmonds, R., Benton, M.J., and Schouten, R.). pdf. Supplementary data file 1; characters, analyses. Supplementary data file 2; PAUP outputs.
Obituary: Dr Arthur Cruickshank, palaeontologist. Herald Scotland, 7th December, 2011 (Taylor, M.A. and Benton, M.J.) Online version.

2010

New take on the Red Queen. Nature 463, 306-307 (doi:10.1038/463306a). pdf.
Fossilized melanosomes and the colour of Cretaceous dinosaurs and birds. Nature 463, 1075-1078 (doi: nature08740.3d) (Zhang, F., Kearns, S.L, Orr, P.J., Benton, M.J., Zhou, Z., Johnson, D., Xu, X., and Wang, X.). pdf. Supplementary data. Interpretive web site.
Phylogenetically structured variance in felid bite force: the role of phylogeny in the evolution of biting performance. Journal of Evolutionary Biology 23, 463-478 (doi:10.1111/j.1420-9101.2009.01922.x) (Sakamoto, M., Lloyd, G.T., and Benton, M.J.). pdf. Featured on journal issue front cover.
Studying function and behavior in the fossil record. PLoS Biology 8(3), e1000321. (doi:10.1371/journal.pbio.1000321). pdf.
The higher-level phylogeny of Archosauria (Tetrapoda: Diapsida). Journal of Systematic Palaeontology 8, 3-47 (Brusatte, S.L., Benton, M.J., Desojo, J.B., and Langer, M.C.. (doi:10.1080/14772010903537732) pdf. Supplementary data.
The origin and radiation of dinosaurs. Earth-Science Reviews 101, 68-100. (doi:10.1016/j.earscirev.2010.04.001) (Brusatte, S.L., Nesbitt, S.J., Irmis, R.B., Butler, R.J., Benton, M.J., and Norell, M.A.). pdf.
The fossil record of early tetrapods: worker effort and the end-Permian mass extinction. Acta Palaeontologica Polonica 55, 213-228 (doi:10.4202/app.2009.002) (Bernard, E.L., Ruta, M., Tarver, J.E. and Benton, M.J.). pdf. Supplementary data.
Disruption of playa-lacustrine depositional systems at the Permo-Triassic boundary: evidence from Vyazniki and Gorokhovets on the Russian Platform. Journal of the Geological Society, London 167, 695-716 (doi: 10.1144/0016-76492009-103) (Newell, A.J., Sennikov, A.G., Benton, M.J., Molostovskaya, I.I., Golubev, V.K., Minikh, A.V., and Minikh, M.G.). pdf. Supplementary data.
An island of dwarfs – reconstructing the Late Cretaceous Hațeg palaeoecosystem. Palaeogeography, Palaeoclimatology, Palaeoecology 293, 265-270 (doi: 10.1016/j.palaeo.2010.05.032) (Csiki, Z. and Benton, M.J). pdf.
Palaeobiogeographic relationships of the Hațeg biota – Between isolation and innovation. Palaeogeography, Palaeoclimatology, Palaeoecology 293, 419-427 (doi: 10.1016/j.palaeo.2010.03.024) (Weishampel, D.B., Csiki, Z., Benton, M.J., Grigorescu, D., and Codrea, V.). pdf.
Dinosaurs and the island rule: The dwarfed dinosaurs from Hațeg Island. Palaeogeography, Palaeoclimatology, Palaeoecology 293, 438-454 (doi:10.1016/j.palaeo.2010.01.026) (Benton, M.J., Csiki, Z., Grigorescu, D., Redelstorff, R., Sander, P.M., Stein, K., and Weishampel, D.B.). pdf.
On the flux ratio method and the number of valid species names. Paleobiology 36, 516-518 (doi:10.1666/09038.1) (Solow, A.R. and Benton, M.J.). pdf.
Links between global taxonomic diversity, ecological diversity and the expansion of vertebrates on land. Biology Letters 6, 544-547 (doi:10.1098/rsbl.2009.1024) (Sahney, S., Benton, M.J., and Ferry, P.A). pdf. Supplementary data. One of the top-ten most downloaded articles from Biology Letters in 2010.
Murchison’s first sighting of the Permian, at Vyazniki in 1841. Proceedings of the Geologists’ Association 121, 313-318 (doi: 10.1016/j.pgeola.2010.03.005) (Benton, M.J., Sennikov, A.G., and Newell, A.J.). pdf.
Diverse tetrapod trackways in the Lower Pennsylvanian Tynemouth Creek Formation, near St. Martins, southern New Brunswick, Canada. Palaeogeography, Palaeoclimatology, Palaeoecology 296, 1-13 (doi:10.1016/j.palaeo.2010.06.020) (Falcon-Lang, H.J., Benton, M.J., Miller, R.F., and Bashforth, A.R.). pdf. Read an interpretive report from Geoscientist.
Recovery of vertebrate faunas from the end-Permian mass extinction. Journal of Earth Science 21, 111-114. pdf.
Naming dinosaur species: the performance of prolific authors. Journal of Vertebrate Paleontology 30, 1478-1485 (doi:10.1080/02724634.2010.501462). pdf. Read a news story in Nature.
The origins of modern biodiversity on land. Philosophical Transactions of the Royal Society, Series B 365, 3667-3679 (doi:10.1098/rstb.2010.0269). pdf.
Rainforest collapse triggered Carboniferous tetrapod diversification in Euramerica. Geology 38, 1079-1082 (doi: 10.1130/G31182.1) (Sahney, S., Benton, M.J., and Falcon-Lang, H.J.). Selected as a Research highlight in Nature, 9th December, 2010. pdf.
Investigating evolutionary radiations. Pp. 10-26. In Long, M. and Zhou, Z. (eds), Darwin’s Heritage Today — Proceedings of the Darwin 200 International Conference. Higher Education Press, Beijing. pdf.
Vertebrate microremains from the Early Cretaceous of southern Tunisia, Geobios 43, 615-628 (doi:10.1016/j.geobios.2010.07.001) (Cuny, G., Cobbett, A.M., Meunier, F.J., Benton, M.J.) pdf.

2009

Introduction to paleobiology and the fossil record. Wiley-Blackwell, Chichester, 592 pp. Further details and book home page.
The Red Queen and the Court Jester: species diversity and the role of biotic and abiotic factors through time. Science 323, 728-732. pdf. Report in El Pais, en español, by Xavier Sampedro.
Paleontology and the history of life. In: Ruse, M. & Travis, J. (eds) Evolution: the first 4 billion years. Harvard Univ. Press, Cambridge, MA, pp. 80-104. ISBN 0-674-03175-X. pdf.
Magnetostratigraphy of Permian/Triassic boundary sequences in the Cis-Urals, Russia: No evidence for a major temporal hiatus. Earth & Planetary Science Letters 281, 36-47 (Taylor, G.K., Tucker, C., Twitchett, R.J., Kearsey, T., Benton, M.J., Newell, A.J., Surkov, M.V., and Tverdokhlebov, V.P.). pdf. Interpretive web site. Press release.
The fossil record: biological or geological signal? In: Sepkoski, D. & Ruse, M. (eds), The paleobiological revolution: essays on the growth of modern paleontology. Univ. Chicago Press, Chicago, 584 pp., pp. 43-59. ISBN 978-0226748610 pdf.
Calibrating and constraining molecular clocks. In: Hedges, S.B. & Kumar, S. (eds) The timetree of life. Oxford University Press, Oxford, pp. 35-86 (Benton, M.J., Donoghue, P.C.J., and Asher, R.J.). pdf. The book web site, with additional resources.
Primer: Dinosaurs. Current Biology 19, R318-R323. pdf.
A reassessment of the Pteraichnus ichnospecies from the Early Cretaceous of Soria Province, Spain. Journal of Vertebrate Paleontology 29, 487-497 (Sánchez-Hernández, B., Przewieslik, A.G., and Benton, M.J.). pdf.
Completeness of the fossil record. Significance 6, 117-121. pdf.
Tetrapod postural shift estimated from Permian and Triassic trackways. Palaeontology 52, 1029-1037 (Kubo, T. and Benton, M.J.). pdf. Interpretive web site.

2008

Palaeoecology of the Late Triassic extinction event in the SW UK. Journal of the Geological Society 165, 319-332 (Mander, L., Twitchett, R.J., and Benton, M.J.). pdf
A new genus of rhynchosaur from the Middle Triassic of southwest England. Palaeontology 51, 95-115 (Hone, D.W.E. and Benton, M.J.). pdf. Read Darren Naish’s very thorough blogs about Fodonyx and other rhynchosaurs here and here, and David Hone’s comments here.
Body size evolution in Mesozoic birds. Journal of Evolutionary Biology 21, 618-624 (Hone, D.W.E., Dyke, G.J., Haden, M. and Benton, M.J.). pdf
When life nearly died: the greatest mass extinction of all time [revised, paperback edition] by M.J. Benton. Book details for people in the UK, and in North America.
Recovery from the most profound mass extinction of all time. Proceedings of the Royal Society, Series B 275, 759-765 (Sahney, S. and Benton, M.J.). pdf
A primitive confuciusornithid bird from China and its implications for early avian flight. Science in China Series D: Earth Sciences 51, 625-639 (Zhang F., Zhou Z., and Benton, M.J.).pdf
The end-Permian mass extinction – events on land in Russia. Proceedings of the Geologists’ Association 119, 119-136. pdf.
The aerodynamics of the British Late Triassic Kuehneosauridae. Palaeontology 51, 967-981 (Stein, K., Palmer, C., Gill, P.G., and Benton, M.J.). pdf
Catastrophic ocean acidification at the Triassic-Jurassic boundary. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 249, 119-127 (Hautmann, M., Benton, M.J., and Tomasovych, A.). pdf
Discussion on ecology of earliest reptiles inferred from basal Pennsylvanian trackways. Journal of the Geological Society, London 165, 985-987 (Falcon-Lang, H. and Benton, M.J.). pdf
Superiority, competition, and opportunism in the evolutionary radiation of dinosaurs. Science 321, 1485-1488 (Brusatte, S.L., Benton, M.J., Ruta, M. and Lloyd, G.T.). pdf. Supplementary data explanation. Interpretive web site.
Dinosaurs and the Cretaceous Terrestrial Revolution. Proceedings of the Royal Society, Series B275, 2483-2490; doi:10.1098/rspb.2008.0715. (Lloyd, G.T., Davis, K.E., Pisani, D., Tarver, J.E., Ruta, M., Sakamoto, M., Hone, D.W.E., Jennings, R., and Benton, M.J.). pdf. Interpretive web site.
(Editor) The seventy great mysteries of the natural world. Thames & Hudson, London, 304 pp. Book details for people in the UK and in North America. Sample text here.
The history of life. A very short introduction. Oxford University Press, Oxford, 170 pp. Book details for people in the UK and in North America.
Fossil quality and naming dinosaurs. Biology Letters 4, 729-732. doi:10.1098/rsbl.2008.0402 pdf. Data supplement. [Reported by Jonathan Amos for BBC News, Will the real dinosaurs stand up, and Rex Dalton for Nature, In search of Thingummyjigosaurus; Holly Williams in The Independent: Under the Microscope: How many types of dinosaur are there?] Interpretive web site.
The first 50 Myr of dinosaur evolution: macroevolutionary pattern and morphological disparity. Biology Letters 4, 733-736. doi:10.1098/rsbl.2008.0441 (Brusatte, S.L., Benton, M.J., Ruta, M. and Lloyd, G.T.). pdf. Interpretive web site.
How to find a dinosaur, and the role of synonymy in biodiversity studies. Paleobiology 34, 516-533. pdf. Interpretive web site.
Calibrated diversity, tree topology and the mother of mass extinctions: the lesson of temnospondyls. Palaeontology 51, 1261-1288 (Ruta, M. and Benton, M.J.). doi:10.1111/j.1475-4983.2008.00808.x pdf.
A new metriorhynchid crocodilian (Mesoeucrocodylia: Thalattosuchia) from the Kimmeridgian (Upper Jurassic) of Wiltshire, UK. Palaeontology 51, 1307-1333 (Wilkinson, L.E., Young, M.T. and Benton, M.J.). doi:10.1111/j.1475-4983.2008.00818.x pdf.
Head kinematics and feeding adaptations of the Permian and Triassic dicynodonts. Journal of Vertebrate Paleontology 28, 1120-1129 (Surkov, M.V. and Benton, M.J.). pdf.
The remarkable fossils from the Early Cretaceous Jehol Biota of China and how they have changed our knowledge of Mesozoic life. Proceedings of the Geologists’ Association 119, 209-228 (Benton, M.J., Zhou Z., Orr, P.J., Zhang F., and Kearns, S.L.). pdf.
Contrasting supertree and total-evidence methods: the origin of the pterosaurs. Zitteliana B28, 35-60 (Hone, D.W.E. and Benton, M.J.). pdf.
Aerodynamic characters of the cranial crest of Pteranodon. Zitteliana 28B, 167-174 (Elgin, R.A., Grau, C.A., Palmer, C., Hone, D.W.E., Greenwell, D., and Benton, M.J.). pdf.

2007

Palaeontological evidence to date the tree of life. Molecular Biology and Evolution 24, 26-53 (Benton, M.J. and Donoghue, P.C.J.). pdf. Designated as an ‘MBE citation classic’ in January 2013, and as a ‘Highly Cited Paper’ by Web of Science.
Palaeoenvironments of vertebrates on the southern shore of Tethys: The nonmarine Early Cretaceous of Tunisia. Palaeogeography, Palaeoclimatology, Palaeoecology 243, 118-131 (Anderson, P.E., Benton, M.J., Trueman, C.N., Paterson, B.A. and Cuny, G.). pdf.
How did life become so diverse? The dynamics of diversification according to the fossil record and molecular phylogenetics. Palaeontology 50, 23-40 (Benton, M.J. and Emerson, B.J.). pdf.
The effects of sampling bias on Palaeozoic faunas and implications for macroevolutionary studies. Palaeontology 50, 177-184 (Tarver, J.E., Braddy, S.J. and Benton, M.J.). pdf.
Modern avian radiation across the Cretaceous-Paleogene boundary. The Auk 124, 339-341 (Dyke, G.J., Nudds, R.L., Benton, M.J.). pdf
Cope’s Rule in the Pterosauria, and differing perceptions of Cope’s Rule at different taxonomic levels. Journal of Evolutionary Biology 20, 1164-1170 (Hone, D.W.E. and Benton, M.J.). pdf
First occurrence of footprints of large therapsids from the Upper Permian of European Russia. Palaeontology 50, 641-652. (Surkov, M. V., Benton, M.J. Twitchett, R. J., Tverdokhlebov, V. P. Newell, A. J. ). pdf.
Dinosaurs and other fossil vertebrates from the Late Jurassic and Early Cretaceous of the Galve area, NE Spain. Palaeogeography, Palaeoclimatology, Palaeoecology 249, 180-215 (Sánchez-Hernández, B., Benton, M.J. and Naish, D.). pdf
Rocks and clocks: calibrating the Tree of Life using fossils and molecules. Trends in Ecology and Evolution 22, 424-431 (Donoghue, P.C.J. and Benton, M.J.). pdf
Bedload abrasion and the in situ fragmentation of bivalve shells. Sedimentology 54, 835-845 (Newell, A. J., Gower, D. J., Benton, M.J., and Tverdokhlebov, V. P.). pdf
Tetrapod tracks from the Mauch Chunk Formation (middle to upper Mississippian) of Pennsylvania, U.S.A. Proceedings of the Academy of Natural Sciences of Philadelphia 156, 199-209 (Vrazo, M. B., Benton, M.J., and Daeschler, E. B.). pdf
The prelude to the end-Permian mass extinction predates a postulated bolide impact. International Journal of Earth Sciences 96, 903-909 (Yin, H.-F., Feng, Q.-l., Baud, A., Xie, S-c., Benton, M.J., Lai, X.-l. and Bottjer, D.J.). pdf
Ecology of earliest reptiles inferred from basal Pennsylvanian trackways. Journal of the Geological Society 164, 1113-1118 (Falcon-Lang, H.J., Benton, M.J., and Stimson, H.). pdf
The Phylocode: beating a dead horse? Acta Palaeontologia Polonica 52, 651-655. pdf.
Congruence of morphological and molecular phylogenies. Acta Biotheoretica 55, 269-281 (Pisani, D., Benton, M.J., and Wilkinson, M.). pdf
A supertree of Temnospondyli: cladogenetic patterns in the most species-rich group of early tetrapods. Proceedings of the Royal Society, Series B 274, 2087-2095 (Ruta, M., Pisani, D., Lloyd, G. T. and Benton, M.J.). pdf.
Evolution of hindlimb posture in archosaurs: limb stresses in extinct vertebrates. Palaeontology 50, 1519-1529 (Kubo, T. and Benton, M.J.). pdf
An evaluation of the phylogenetic relationships of the pterosaurs among archosauromorph reptiles. Journal of Systematic Palaeontology 5, 465-469 (Hone, D.W.E. and Benton, M.J.). pdf

2006

The Pennsylvanian tropical biome reconstructed from the Joggins Formation of Nova Scotia, Canada. Journal of the Geological Society, London 163, 561-576 Falcon-Lang, H.J., Benton, M.J., Braddy, S.J. and Davies, S.J. pdf
Pelagosaurus typus Bronn, 1841 (Mesoeucrocodylia: Thalattosuchia) from the Upper Lias (Toarcian, Lower Jurassic) of Somerset, England. Journal of Vertebrate Paleontology 26, 621-635 (S. Pierce & Benton, M.J.). pdf
The origin of the dinosaurs. In III Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno, pp. 11-19. Colectivo Arqueológico-Paleontológico de Salas, Burgos, Spain. pdf.
Early dinosaurs: a phylogenetic study. Journal of Systematic Palaeontology 4, 309-358 (M. C. Langer & Benton, M.J.). pdf.
Dwarfing in ornithopod dinosaurs from the Early Cretaceous of Romania. In: Csiki, Z. (ed.) Mesozoic and Cenozoic Vertebrates and Paleoenvironments; Tributes to the Career of Prof. Dan Grigorescu, Ars Docendi, Bucharest, 79-87 (Benton, M.J., N. J. Minter and E. Posmosanu). pdf

2005

The evolution of large size: how does Cope’s Rule work? Trends in Ecology and Evolution 20, 4-6 (Hone, D.W.E. and Benton, M.J.). pdf
Upper Permian vertebrates and their sedimentological context in the South Urals, Russia. Earth-Science Reviews, 69, 27-77 (Tverdokhlebov, V.P., Tverdokhlebova, G.I., Minikh, A.V., Surkov, M.V., and Benton, M.J.). pdf.
The quality of the fossil record of Mesozoic birds. Proceedings of the Royal Society, Series B 272, 289-294 (Fountaine, T.M.R., Benton, M.J., Dyke, G.J. and Nudds, R.L.). pdf The data may be downloaded here.
Lystrosaurus georgi, a dicynodont from the Lower Triassic of Russia. Journal of Vertebrate Paleontology 25, 402-413 (Surkov, M. V., Kalandadze, N. N. and Benton, M.J.). pdf
Fossil record. Encyclopedia of life sciences. Macmillan, London, 11 pp. pdf
Fossil record: quality. Encyclopedia of life sciences. Macmillan, London, 11 pp. pdf
Reptilia. Encyclopedia of life sciences. Macmillan, London, 11 pp. pdf
Sauria. Encyclopedia of life sciences. Macmillan, London, 9 pp. pdf
Occurrence of sauropod dinosaur tracks in the Upper Jurassic of Chile (redescription of Iguanodonichnus frenki). Journal of South American Earth Sciences 20, 253-257 (Moreno, K. and Benton, M.J.). pdf
Mesozoic and Tertiary fossil mammals and birds of Great Britain. Joint Nature Conservancy Committee, London, 215 pp. (Benton, M.J., E. Cook and J. J. Hooker). Downloads.

2004

The basicranium of dicynodonts (Synapsida) and its use in phylogenetic systematics. Palaeontology 47, 619-638 (Surkov, M.V. and Benton, M.J.). pdf
Ecosystem remodelling among vertebrates at the Permian-Triassic boundary in Russia. Nature 432, 97-100 (doi:10.1038/nature02950) (Benton, M.J., Tverdokhlebov, V.P. and Surkov, M.V.). pdf Download the original of Figure 1 in colour as a pdf. Download the Excel data file here.
Origin and relationships of Dinosauria. In D. B. Weishampel, P. Dodson, and H. Osmólska (eds), The Dinosauria, 2nd edition, pp. 7-24. University of California Press, Berkeley. pdf
Neoselachian (Chondrichthyes, Elasmobranchii) diversity across the Cretaceous-Tertiary boundary. Palaeogeography, Palaeoclimatology, Palaeoecology 214, 181-194 (Kriwet, J. and Benton, M.J.). pdf
Fossil sharks from the Early Cretaceous of Tunisia. Revue de Paléontologie, Volume spéciale 9, 127-142 (Cuny, G., Ouaja, M., Srarfi, D., Schmitz, L., Buffetaut, E., and Benton, M.J.). pdf

2003

Tetrapod localities from the Triassic of the SE of European Russia. Earth-Science Reviews 60, 1-66 (Tverdokhlebov, V.P., Tverdokhlebova, G.I., Surkov, M.V., and Benton, M.J.). pdf.
When life nearly died: the greatest mass extinction of all time. Thames & Hudson, London, New York, 336 pp. Read more about this book.
Wipeout. New Scientist 178 (2392), 38-41. pdf version of MS
Dating the tree of life. Science 300, 1698-1700 (Benton, M.J. and Ayala, F.J.). pdf. Full copy of the supplementary material.
How to kill (almost) all life: the end-Permian extinction event. Trends in Ecology and Evolution 18, 358-365 (doi:10.1016/S0169-5347(03)00093-4) (Benton, M.J. and Twitchett, R.J.). pdf
Geochemical taphonomy of shallow marine vertebrate assemblages. Palaeogeography, Palaeoecology, Palaeoclimatology 197, 151-169 (Trueman, C. N., Benton, M.J., and Palmer, M.R.). pdf
Post-Cambrian closure of the deep-water slope-basin taphonomic window. Geology 31, 769-772 (Orr, P. J., Benton, M.J., and Briggs, D. E. G.). pdf
Taxonomic level as a determinant of the shape of the Phanerozoic marine biodiversity curve. American Naturalist 162, 265-276 (Lane, A. and Benton, M.J.). pdf
The quality of the fossil record. Pp. 66-90 in Donoghue, P. C. J. and Smith, M. P. (eds) Telling the Evolutionary Time: Molecular Clocks and the Fossil Record. Taylor & Francis, London. pdf
Patterns and rates of species evolution. Encyclopedia of Life Support Systems. pdf

2002

Permian and Triassic red beds and the Penarth Group of Great Britain. Joint Nature Conservation Committee, Peterborough, xvi + 337 pp. (Benton, M.J., E. Cook, and P. Turner). Text downloads and image downloads. pdf of Chapter 4, Penarth Group.
A genus-level supertree of the Dinosauria. Proceedings of the Royal Society, Series B, 269, 915-921. (Pisani, D., Yates, A.M., Langer, M.C., and Benton, M.J.). pdf. Interpretive web site.
Cope’s Rule. Pp. 209-210 in Pagel, M. (ed.) Encyclopedia of evolution. Oxford University Press, New York. pdf
Alick Walker 1925-1999: an appreciation. Zoological Journal of the Linnean Society 136, 1-5 (Benton, M.J. and Gower, D.J.). pdf.
Erpetosuchus, a crocodile-like basal archosaur from the Late Triassic of Elgin, Scotland. Zoological Journal of the Linnean Society 136, 25-47 (Benton, M.J. and Walker, A.D.). pdf.

2001

Evidence of evolutionary transitions. ActionBioscience
Longisquama fossil and feather morphology. Science 291, 1900-1901 (Unwin, D. M. and Benton, M.J.). pdf.
Biodiversity through time. Pp. 211-220 in Briggs, D. E. G. and Crowther, P. R. (eds) Palaeobiology II. Blackwell Science, Oxford. pdf
Speciation in the fossil record. Trends in Ecology and Evolution 16, 405-411 (Benton, M.J. and P. N. Pearson). pdf
The science of ‘Walking with dinosaurs’. Teaching Earth Sciences 26, 4-9. pdf pdf of MS.
Biodiversity on land and in the sea. Geological Journal 36, 211-230. pdf
Finding the tree of life: matching phylogenetic trees to the fossil record through the 20th century. Proceedings of the Royal Society of London, Series B 268, 2123-2130. pdf

2000

Quality of the fossil record through time. Nature 403, 534-538 (Benton, M.J., Wills, M. and Hitchin, R.). pdf.
Dinosaurs and other fossil vertebrates from fluvial deposits in the Lower Cretaceous of southern Tunisia. Palaeogeography, Palaeoclimatology, Palaeoecology 157, 227-246 (Benton, M.J., S. Bouaziz, E. Buffetaut, D. Martill, M. Ouaja, M. Soussi, and C. Trueman). pdf
Anatomy and systematics of the prosauropod dinosaur Thecodontosaurus antiquus from the Upper Triassic of southwest England. Journal of Vertebrate Palaeontology 20, 77-108 (Benton, M.J., L. Juul, G. W. Storrs, and P. M. Galton). pdf
The fossil record of Cretaceous tetrapods. Palaios 15, 161-165 (E. Fara and Benton, M.J.). pdf
Stems, nodes, crown clades, and rank-free lists: is Linnaeus dead? Biological Reviews 75, 633-648. pdf
The age of dinosaurs in Russia and Mongolia (eds Benton, M.J., M. A. Shishkin, D. M. Unwin, and E. N. Kurochkin. Cambridge University Press, Cambridge, 696 pp. Read the scanned book at Google Books.
Conventions in Russian and Mongolian palaeontological literature. Pp. xvi-xxxix, in Benton, M. J., Shishkin, M. A., Unwin, D. M., and Kurochkin, E. N. (Eds) The age of dinosaurs in Russia and Mongolia. Cambridge University Press, Cambridge. pdf
Procolophonoids from the Permo-Triassic of Russia. Pp. 160-176, in Benton, M. J., Shishkin, M. A., Unwin, D. M., and Kurochkin, E. N. (Eds) The age of dinosaurs in Russia and Mongolia. Cambridge University Press, Cambridge (Spencer, P. S. and Benton, M.J.). pdf
Enigmatic small reptiles from the Middle-Late Triassic of Kirgizstan. Pp. 177-186, in Benton, M. J., Shishkin, M. A., Unwin, D. M., and Kurochkin, E. N. (Eds) The age of dinosaurs in Russia and Mongolia. Cambridge University Press, Cambridge (Unwin, D. M., Alifanov, V. R., and Benton, M.J.). pdf
A brief history of dinosaur paleontology. Pp. 10-44, in Paul, G. S. (ed.), The Scientific American book of dinosaurs. St Martin’s Press, New York. pdf of text.
Accuracy of fossils and dating techniques. ActionBioscience
Crown clades in vertebrate nomenclature: correcting the definition of Crocodylia. Macroevolution. Pp. 155-166, in Lane, R. H., Steininger, F. F., Kaesler, R. L., Ziegler, W., and Lipps, J. (eds), Fossils and the future: paleontology in the 21st century. Senckenberg-Buch Nr. 74, Frankfurt-am-Main (Jablonski, J., Benton, M.J., Gastaldo, R. A., Marshall, C. R., and Sepkoski, J. J., Jr.). html pdf
Mammals. Pp. 639-644, in The Oxford Companion to the Earth, P. L. Hancock and B. J. Skinner (eds), Oxford University Press, 1174 pp. pdf.
Discussion on Lazarus taxa and fossil abundance at times of biotic crisis. Journal of the Geological Society, London 157, 512 (P. B. Wignall and Benton, M.J.).
Criticality and scaling in evolutionary biology. Trends in Ecology and Evolution 14, 156-160 (Solé, R. V., Manrubia, S. C., Benton, M. J., Kauffman, S., and Bak, P.). pdf

1999

Evolutionary patterns from mass originations and mass extinctions. Proceedings of the Royal Society of London, Series B 354, 463-469 (Hewzulla, D., Boulter, M.C., Benton, M.J., and Halley, J.M.). pdf
Lazarus taxa and abundance at times of biotic crisis. Journal of the Geological Society of London 156, 453-456 (P. Wignall and Benton, M.J.). pdf
The history of life: large databases in palaeontology. In D. A. T. Harper (ed.), Numerical palaeobiology. Wiley, Chichester, pp. 249-283. pdf
Early radiation of the neoselachian sharks in western Europe. Geobios 32, 193-204 (G. Cuny and Benton, M.J.). pdf
Self-organized criticality in ecology and evolution. Trends in Ecology and Evolution 14, 321-322 (Solé, R.V., Manrubia, S. C., Benton, M.J., Kauffman, S., and Bak, P.) pdf
Scleromochlus taylori and the origin of dinosaurs and pterosaurs. Philosophical Transactions of the Royal Society of London, Series B 354, 1423-1446. pdf.
Interplay of tectonics and climate on a transverse fluvial system, Upper Permian, southern Uralian foreland basin, Russia. Sedimentary Geology 127, 11-29 (Newell, A. J., Tverdokhlebov, V. P., and Benton, M.J.). pdf
A sauropodomorph dinosaur from the Upper Triassic (Carnian) of southern Brazil. Comptes rendus de l’Academie des Sciences, Paris, Sciences de la Terre et des Planètes 329, 511-517 (Langer, M. A., Abdala, F., Richter, M., and Benton, M.J.). pdf
Assessing congruence between cladistic and stratigraphic data. Systematic Biology 48, 581-596 (Benton, M.J., R. Hitchin, and M. A. Wills). pdf
Why I dared walk with TV dinosaurs. The Times November 12, 1999, p. 17. MS copy of article.
Early origins of modern birds and mammals: molecules vs. morphology. BioEssays 21, 1043-1051 (and Reply to Easteal. BioEssays 21, 1059). pdf

1998

Analysing diversification through time: reply to Sepkoski and Miller. Trends in Ecology and Evolution 13, 201. pdf
Dinosaur fossils with soft parts. Trends in Ecology and Evolution 13, 303-304. pdf
The quality of the fossil record of vertebrates. Pp. 269-303, in Donovan, S. K. and Paul, C. R. C. (eds), The adequacy of the fossil record. Wiley, New York, 312 pp. MS pdf version
Molecular and morphological phylogenies of mammals: congruence with stratigraphic data. Molecular Phylogenetics and Evolution 9, 398-407. pdf
Robust dinosaur phylogeny? Nature 396, 423-424 (Wilkinson, M., Upchurch, P., Barrett, P. M., Gower, D. J., and Benton, M.J.). pdf
Long-range correlations in the fossil record and the fractal nature of macroevolution. Advances in Complex Systems 1, 255-266 (Solé, R. V., Manrubia, S.C., Peerez-Mercader, J., Benton, M.J., and Bak, P.). pdf
Palaeoenvironmental interpretation of the Triassic sandstones of Scrabo, County Down, Northern Ireland: ichnological and sedimentological studies indicating a mixed fluviatile-aeolian succession. Irish Journal of Earth Sciences 16, 85-102 (Buckman, J. O., Doughty, P. S., Benton, M.J., and Jeram, A. J.). pdf

1997

First record of footprints of terrestrial vertebrates from the Upper Permian of the Cis-Urals, Russia. Palaeontology 40, 157-166 (Tverdokhlebov, V. P., Tverdokhlebova, G. I., Benton, M.J., and Storrs, G. W.). pdf
A geochemical method to trace the taphonomic history of reworked bones in sedimentary settings. Geology, 25, 263-266 (C. N. Trueman and Benton, M.J.). pdf
Uncertain turtle relationships. Nature, 387, 466 (M. Wilkinson, J. Thorley, and Benton, M.J.). pdf
Congruence between parsimony and stratigraphy: comparisons of three indices. Paleobiology 23, 20-32 (R. Hitchin and Benton, M.J.). pdf
Richard Owen’s giant Triassic frogs: archosaurs from the Middle Triassic of England. Journal of Vertebrate Paleontology 17, 74-88 (Benton, M.J. and D. J. Gower). pdf
Dinosaurs and other tetrapods in an Early Cretaceous bauxite-filled fissure, northwestern Romania. Palaeogeography, Palaeoclimatology, Palaeoecology 130, 275-292 (Benton, M.J., Cook, E., Grigorescu, D., Popa, E., and Tallodi, E.). pdf
Congruence between phylogenetic and stratigraphic data on the history of life. Proceedings of the Royal Society of London, Series B 264, 885-890 (Benton, M.J. and R. Hitchin). pdf
Self-similarity of extinction statistics in the fossil record. Nature 388, 764-767 (R. V. Solé, S. C. Manrubia, Benton, M.J., and P. Bak). pdf
Origin and early evolution of dinosaurs. Pp. 204-215 In J. O. Farlow and M. K. Brett-Surman (eds), The complete dinosaur. Indiana University Press, Bloomington, Indiana, 752 pp. Read this chapter at Google Books
Stratigraphic indices and tree balance. Systematic Biology 46, 563-569 (R. Hitchin and Benton, M.J.). pdf
Models for the diversification of life. Trends in Ecology and Evolution 12, 490-495. pdf
Boreopricea from the Lower Triassic of Russia, and the relationships of the prolacertiform reptiles. Palaeontology 40, 931-953 (Benton, M.J. and J. Allen). pdf
Palaeoenvironmental interpretation of the Triassic Sandstone of Scrabo, County Down, Northern Ireland: ichnological and sedimentological studies indicating a mixed fluviatile-aeolian succession. Irish Journal of Earth Sciences 16, 85-102 (Buckman, J.O., Doughty, P.S., Benton, M.J., Jeram, A.J.). pdf
The Triassic reptiles from Devon. Proceedings of the Ussher Society 9, 141-152. pdf

1996

Diversity in the past: comparing cladistic phylogenies and stratigraphy. Pp. 19-40. In Hochberg, M. E., Clobert, J., and Barbault, R. (eds) Aspects of the genesis and maintenance of biological diversity. Oxford University Press, Oxford, Oxford, 316 pp. (Benton, M.J. and Storrs, G. W.). pdf
Testing the marine and continental fossil records: Reply. Geology, 23, 381-382. pdf
Lower Silurian trace fossils and the Eocoelia community in the Tortworth Inlier, SW England. Proceedings of the Geologists’ Association 107, 199-208 (Benton, M.J. and C. Hiscock). pdf
Testing the roles of competition and expansion in tetrapod evolution. Proceedings of the Royal Society of London, Series B 263, 641-646. pdf
Kimmeridgian metriorhynchid crocodiles from England. Palaeontology 39, 497-514 (D. R. Grange and Benton, M.J.). pdf
Dinosaurs in the Early and Mid Triassic? – The footprint evidence from Britain. Palaeogeography, Palaeoclimatology, Palaeoecology 122, 213-225 (M. J. King and Benton, M.J.). pdf
Rhombopholis, a prolacertiform reptile from the Middle Triassic of England. Palaeontology 39, 763-782 (Benton, M.J. and A. D. Walker). pdf
Deep marine trace fossil assemblages from the Lower Carboniferous of Menorca, Balearic Islands, western Mediterranean. Geological Journal 31, 235-258 (P. J. Orr, Benton, M.J., and N. H. Trewin). pdf
Testing the quality of the fossil record by groups and by major habitats. Historical Biology 12, 111-157 (Benton, M.J. and R. Hitchin). pdf
On the nonprevalence of competitive replacement in the evolution of tetrapods. Pp. 185-210. In Jablonski, D., Erwin, D. H., and Lipps, J. H. (eds) Evolutionary paleobiology. University of Chicago Press, Chicago, 484 pp. pdf

1995

Reply from M. J. Benton [to comment by M. A. Norell on 1994]. Trends in Ecology and Evolution 10, 37. pdf
Diversification and extinction in the history of life. Science, 268, 52-58. pdf
Fossil Reptiles of Great Britain. Chapman and Hall, London, 386 pp. (Benton, M.J. and Spencer, P.S.). Text downloads, and image downloads. The whole book as a single pdf.
Shelf storm beds on the southern margin of the Welsh Basin: the Wenlock of the Tortworth Inlier, England. Proceedings of the Geologists’ Association 106, 81-92 (Benton, M.J. and Thackray, E.). pdf
Testing the time axis of phylogenies. Philosophical Transactions of the Royal Society, Series B, 349, 5-10. pdf
Early Jurassic mass extinction: a global long-term event. Geology, 23, 495-498 (Little, C.T.S. and Benton, M.J.). pdf
Testing the marine and continental fossil records. Geology, 23, 601-604 (Benton, M.J. and Simms, M.J.). pdf
Mass extinctions and periodicity. Reply. Science, 269, 618-619. pdf
Missing data and rhynchosaur phylogeny. Historical Biology 10, 137-250. (Wilkinson, M. and Benton, M.J.). pdf
The first Lower Jurassic dinosaur from Scotland: limb bone of a ceratosaur theropod from Skye. Scottish Journal of Geology 31, 177-182 (Benton, M.J., Martill, D.M., and Taylor, M.A.). pdf

1994

Testing the quality of the fossil record: paleontological knowledge is improving. Geology, 22, 111-114 (Benton, M.J. and Storrs, G.W.). pdf.
Late Triassic terrestrial vertebrate extinctions: stratigraphic aspects and the record of the Germanic Basin. Paleontologia Lombarda, Nuova Serie 2, 19-38. pdf.
Palaeontological data and identifying mass extinctions. Trends in Ecology and Evolution 9, 181-185. pdf
Professor R. J. G. Savage: an appreciation. Zoological Journal of the Linnean Society 112, 3-12. pdf.
A review of the British Middle Triassic tetrapod assemblages. In Fraser, N. C. and Sues, H.-D. (eds), In the shadow of the dinosaurs, pp. 131-160. Cambridge University Press, Cambridge, 435 pp. (Benton, M.J., Warrington, G., Newell, A.J., and Spencer, P.S.). pdf.
Late Triassic to Middle Jurassic extinctions among continental tetrapods: testing the pattern. In Fraser, N. C. and Sues, H.-D. (eds), In the shadow of the dinosaurs, pp. 366-397. Cambridge University Press, Cambridge, 435 pp. pdf
Graptolite evidence for the age of the Dunquin Group (Silurian), Dingle Peninsula, County Kerry. Irish Journal of Earth Sciences 13, 91-94 (Benton, M.J. and Underwood, C.J.). pdf.
Impact in the Caribbean and death of the dinosaurs. Geology Today 10, 222-227 (Benton, M.J. and Little, C.T.S.). pdf.

1993

A new Bathonian (Middle Jurassic) microvertebrate site, within the Chipping Norton Limestone Formation at Hornsleasow Quarry, Gloucestershire. Proceedings of the Geologists’ Association 103, 321-342 (Metcalf, S. J., Vaughan, R. F., Benton, M.J., Cole, J., Simms, M. J. and Dartnall, D. L.) pdf
Late Triassic extinctions and the origin of the dinosaurs. Science, 260, 769-770. pdf
A new rhynchosaur from the middle Triassic of Devon. Proceedings of the Ussher Society 8, 167-171 (Benton, M.J., M. B. Hart, and T. Clarey). pdf
Preface. In Benton, M. J. (ed.) The fossil record 2, pp. xi-xvii. Chapman and Hall, London, 845 pp. pdf
Mollusca: Amphineura and ‘Monoplacophora’. In Benton, M. J. (ed.) The fossil record 2, pp. 125-130. Chapman and Hall, London, 845 pp (Benton, M.J. and D. H. Erwin).
Mollusca: Rostroconchia, Scaphopoda and Bivalvia. In Benton, M. J. (ed.) The fossil record 2, pp. 237-263. Chapman and Hall, London, 845 pp (Skelton, P. W. and Benton, M.J.).
Basal deuterostomes (chaetognaths, hemichordates, calcichordates, cephalochordates and tunicates. In Benton, M. J. (ed.) The fossil record 2, pp. 529-535. Chapman and Hall, London, 845 pp.
Reptilia. In Benton, M. J. (ed.) The fossil record 2, pp. 681-715. Chapman and Hall, London, 845 pp. pdf

1992

Nothing. Must buck up.

1991

Polar dinosaurs and ancient climates. Trends in Ecology and Evolution 6, 28-30. pdf
The myth of the Mesozoic cannibals. New Scientist 132 (1790), 40-44. pdf
What really happened in the Late Triassic? Historical Biology 5, 263-278. pdf
Amniote phylogeny. In Trueb, L. and Schultze, H.-P. (eds) Origins of the higher groups of tetrapods: controversy and consensus, pp. 317-330. Cornell University Press, Ithaca, 720 pp. pdf
Extinction, biotic replacements, and clade interactions. Pp. 78-102. In E. Dudley (Ed.) The Unity of Evolutionary Biology. Dioscorides Press, Portland, Oregon, 1160 pp. pdf
Why is life so diverse? In Dudley, E. (ed.) The unity of evolutionary biology, pp. 255-261. Dioscorides Press, Portland, Oregon, 1160 pp. pdf

1990

Scientific methodologies in collision: the history of the study of the extinction of the dinosaurs. Evolutionary Biology, 24, 371-424. pdf of photocopy. pdf of seachable scan.
Phylogeny of the major tetrapod groups: morphological data and divergence dates. Journal of Molecular Evolution 30, 409-424. pdf
Mass extinctions in the fossil record of late Palaeozoic and Mesozoic tetrapods. In Kauffman, E.G. and Walliser, O.H. (eds) Extinction events in earth history; Lecture Notes in the Earth Sciences 30, 239-251. Springer, Berlin. (doi:10.1007/BFb0011129) pdf
The species of Rhynchosaurus, a rhynchosaur (Reptilia, Diapsida) from the Middle Triassic of England. Philosophical Transactions of the Royal Society, London, Series B, 328, 213-306. pdf
Reptiles. In K. McNamara (ed.) Evolutionary trends, pp. 279-300. Belhaven, London. pdf
Origin and interrelationships of dinosaurs. In D. B. Weishampel, P. Dodson, and H. Osmólska (eds), The Dinosauria, pp. 11-30. University of California Press, Berkeley. pdf
Phylogenetic trees and the unification of systematic biology. Trends in Ecology and Evolution 5, 393-394. pdf
The causes of the diversification of life. In P. D. Taylor and G. Larwood (eds), Major evolutionary radiations. Systematics Association Special Volume 42, 409-430. Clarendon, Oxford. pdf
Historical tests of the absolute completeness of the fossil record of tetrapods. Paleobiology 16, 322-335 (W. D. Maxwell and Benton, M.J.). pdf

1989

Fossil reptiles from ancient caves. Nature 337, 309-310. pdf
Patterns of evolution and extinction in vertebrates. In Allen, K. C. and Briggs, D. E. G. (eds.) Evolution and the fossil record. Belhaven, London, pp. 218-241. pdf
Heterochrony in a fossil reptile: juveniles of the rhynchosaur Scaphonyx fischeri from the Late Triassic of Brazil. Palaeontology 32, 335-353. (Benton, M.J. and R. Kirkpatrick). pdf
The Triassic reptiles Brachyrhinodon and Polysphenodon and the relationships of the sphenodontids. Zoological Journal of the LInnean Society 96, 413-445. (N. C. Fraser and Benton, M.J.). doi:0.1111/j.1096-3642.1989.tb02521.x pdf
Pruning the tree of life. Nature 342, 129-130. pdf
The future of earth sciences site conservation in Great Britain. Geological Curator 5, 101-109. (P. Allen, MJB, G. P. Black, C. J. Cleal, K. M. Evans, S. I. Jusypiw, M. A. Rowlands, and T. S. Westoll.) pdf.
Mass extinctions among tetrapods and the quality of the fossil record. Philosophical Transactions of the Royal Society, London, Series B, 325, 369-386. pdf
Red Queen hypothesis. In Briggs, D. E. G. and Crowther, P. R. (eds), Palaeobiology, a synthesis, pp. 119-124. Blackwells, Oxford. pdf
Evolution of large size. In Briggs, D. E. G. and Crowther, P. R. (eds), Palaeobiology, a synthesis, pp. 147-152. Blackwells, Oxford. pdf
End-Triassic. In Briggs, D. E. G. and Crowther, P. R. (eds), Palaeobiology, a synthesis, pp. 194-198. Blackwells, Oxford. pdf

1988

Burrowing by vertebrates. Nature 331, 17-18. pdf
Joints of the crocodile-reversed archosaurs (A. R. I. Cruickshank and Benton, M.J.). Nature 331, 218. pdf
The relationships of the major groups of mammals: new approaches. Trends in Ecology and Evolution 2, 40-45. pdf
The nature of an adaptive radiation. Trends in Ecology and Evolution 3, 127-128. pdf
Mass extinctions in the fossil record of reptiles: paraphyly, patchiness, and periodicity(?) In Larwood, G. (ed.) Extinction and survival in the fossil record. Systematics Association Special Volume 34, 269-294. pdf
The origins of the dinosaurs. Modern Geology 13, 41-56. pdf
Bringing up baby. Nature 334, 566. pdf
Archosaur phylogeny and the relationships of the Crocodylia. In Benton, M. J. (ed.) The phylogeny and classification of the tetrapods. Systematics Association Special Volume, 35A, 295-338 (Benton, M.J. and J. Clark). pdf Supplementary data – data matrix used in generating basal archosaur cladograms.
British fossil reptile sites. In Crowther, P. and Wimbledon, W. A. (eds.) The use and conservation of palaeontological sites. Special Paper in Palaeontology 40, 73-84. pdf

1987

The mite pockets of lizards. Nature 325, 391-392. pdf
Conodonts classified at last. Nature 325, 482-483. pdf
Trace fossils from the Rhaetic shore-face deposits of Staffordshire. Palaeontology 30, 407-428 (A. D. Wright and Benton, M.J.). pdf
The history of the biosphere: equilibrium and non-equilibrium models of global diversity. Trends in Ecology and Evolution 2, 153-156. pdf
The phylogeny of rhynchosaurs (Reptilia; Diapsida; Triassic), and two new species. In P. J. Currie and E. Koster (eds), Fourth symposium on Mesozoic terrestrial ecosystems, Short papers. Tyrrell Museum of Palaeontology, Occasional Papers, 3, 12-17. pdf
Mass extinctions and data bases: changes in the interpretations of tetrapod mass extinctions in the past 20 years. In P. J. Currie and E. Koster (eds), Fourth symposium on Mesozoic terrestrial ecosystems, Short papers. Tyrrell Museum of Palaeontology, Occasional Papers, 3, 156-160. pdf
Progress and competition in macroevolution. Biological Reviews 62, 305-338. pdf
Mass extinctions among families of non-marine tetrapods: the data. Mémoires de la Société Géologique de France 150, 21-32. pdf

1986

Predation by drilling gastropods. Nature 321, 110-111. pdf
Sedimentological uses of dinosaurs. Nature 321, 732. pdf
More than one event in the late Triassic mass extinction. Nature 321, 857-861. pdf
Diapsida Permo-Triadica. Fossilium Catalogus 128, 1-50. Kugler Publications, Amsterdam.
The late Triassic reptile Teratosaurus – a rauisuchian, not a dinosaur. Palaeontology 29, 293-301. pdf
Reptiles from the Upper Kimmeridge Clay (Kimmeridgian, Upper Jurassic) of the vicinity of Egmont Bight. Proceedings of the Dorset Natural History and Archaeological Society 107, 121-125 (M. A. Taylor and Benton, M.J.). pdf
The demise of a living fossil. Nature 323, 762. pdf
The evolutionary significance of mass extinctions. Trends in Ecology and Evolution 1, 127-130. pdf
The Late Triassic tetrapod extinction events. In Padian, K. (ed.), The beginning of the age of dinosaurs; faunal change across the Triassic-Jurassic boundary, pp. 303-320. Cambridge Univ. Press. pdf.

1985

The conservation and use of fossil vertebrate sites: British fossil reptile sites. Proceedings of the Geologists’ Association 96, 1-6 (Benton, M.J. and Wimbledon, W.A.). pdf
To clone a dinosaur. New Scientist 105 (1439), 41-43. pdf
The Red Queen put to the test. Nature 313, 734-735. pdf
Interpretations of mass extinction. Nature 314, 496-497. pdf
Palaeoecology, taphonomy and dating of Permo-Triassic reptiles from Elgin, north-east Scotland. Palaeontology 28, 207-234 (Benton, M.J. and A. D. Walker). pdf
Classification and phylogeny of the diapsid reptiles. Zoological Journal of the Linnean Society 84, 97-164. pdf
Mass extinctions. Nature 315, 536. pdf
Patterns in the diversification of Mesozoic non-marine tetrapods and problems in historical diversity analysis. Special Papers in Palaeontology 33, 185-202. pdf
Mass extinction among non-marine tetrapods. Nature 316, 811-814. pdf
Dinosaurs that fill the gaps. Nature 317, 199. pdf
Archosaur ankles and the relationships of the thecodontian and dinosaurian reptiles. Nature 317, 715-717 (A. R. I. Cruickshank and Benton, M.J.). pdf
The first marsupial fossil from Asia. Nature 318, 313. pdf

1984

Small companions for early dinosaurs. Nature 307, 111-112. pdf
Marine reptiles from the Upper Lias (Lower Toarcian, Lower Jurassic) of the Yorkshire coast. Proceedings of the Yorkshire Geological Society 44, 399-429 (Benton, M.J. and M. A. Taylor). pdf
Dinosaurs’ lucky break. Natural History 93 (6), 54-59. pdf
Rauisuchians and the success of archosaurs. Nature 310, 101. pdf
Fossil reptiles of the German late Triassic and the origin of the dinosaurs. In: W.-E. Reif and F. Westphal (eds), Third symposium on Mesozoic terrestrial ecosystems, Tübingen, Short papers, pp. 13-18. Attempto, Tübingen. pdf
Giant tortoises down under. Nature 311, 303. pdf
The relationships and early evolution of the Diapsida. Symposium of the Zoological Society of London 52, 575-596. pdf
Consensus on archosaurs. Nature 312, 599. pdf
Tooth form, growth and function in Triassic rhynchosaurs (Reptilia, Diapsida). Palaeontology 27, 737-776. pdf

1983

Large-scale replacements in the history of life. Nature 302, 16-17. pdf
Dinosaur success in the Triassic: a noncompetitive ecological model. Quarterly Review of Biology 58, 29-55. pdf
The age of the rhynchosaur. New Scientist 98, 9-13. pdf
Progressionism in the 1850s: Lyell, Owen, Mantell and the Elgin fossil reptile Leptopleuron (Telerpeton). Archives of Natural History 11, 123-136. pdf
Paleozoic trace fossils from the Kufrah Basin, Libya. Journal of Paleontology 57, 447-460 (B. R. Turner and Benton, M.J.). pdf
No consensus on Archaeopteryx. Nature 305, 99-100. pdf
The Triassic reptile Hyperodapedon from Elgin: functional morphology and relationships. Philosophical Transactions of the Royal Society of London, Series B, 302, 605-717. pdf

1982

The Diapsida: revolution in reptile relationships. Nature 296, 306-307. pdf
Multidirectional palaeocurrents as indicators of shelf storm beds. In G. Einsele and A. Seilacher (eds.), Cyclic and event stratification, pp. 350-353. Springer Verlag, Berlin (D. I. Gray and Benton, M.J.). pdf
Dictyodora and associated trace fossils from the Palaeozoic of Thuringia. Lethaia 15, 115-132. pdf.
Trace fossils from the Lower Palaeozoic ocean-floor sediments of the Southern Uplands of Scotland. Transactions of the Royal Society of Edinburgh: Earth Sciences 73, 67-87. pdf
Triassic environments, climates and reptile evolution. Palaeogeography, Palaeoclimatology, Palaeoecology 40, 361-379 (M. E. Tucker and Benton, M.J.). pdf

1981

The use of flexible synthetic rubbers for casts of complex fossils from natural moulds. Geological Magazine 118, 551-556 (Benton, M.J. and A. D. Walker). pdf
Lower Silurian distal shelf storm-induced turbidites in the Welsh Borders: sediments, tool marks and trace fossils. Journal of the Geological Society, London 138, 675-694 (Benton, M.J. and D. I. Gray). pdf

1980

H. A. Nicholson (1844-1899), invertebrate palaeontologist: bibliography and catalogue of his type and figured material. Royal Scottish Museum Information Series, Geology 7, 1-94. pdf.
Geographic variation in the garter snakes (Thamnophis sirtalis) of the north-central United States, a multivariate study. Zoological Journal of the Linnean Society 68, 307-323. doi:10.1111/j.1096-3642.1980.tb01924.x pdf
Dictyodora from the Silurian of Peeblesshire, Scotland. Palaeontology 23, 501-513 (Benton, M.J. and N. H. Trewin). pdf
Geographic variation and the validity of subspecies names for the Eastern garter snake, Thamnophis sirtalis. Bulletin of the Herpetological Society of Chicago 15, 57-69. pdf

1979

Ecological succession among late Palaeozoic and Mesozoic tetrapods. Palaeogeography, Palaeoclimatology, Palaeoecology 26, 127-150. pdf
Increase in total global biomass over time. Evolutionary Theory 4, 123-128. pdf
Ectothermy and the success of the dinosaurs. Evolution 33, 983-997. pdf
H. A. Nicholson (1844-1899): pioneer of thin-section taxonomy. Fossil Cnidaria 8 (2), 17-24. pdf

1978

Discussion and comments on Nicholson’s 1872 manuscript ‘Contributions to the study of errant annelides of the older Palaeozoic rocks’. Publications of the Department of Geology and Mineralogy, University of Aberdeen 1, 1-16 (Benton, M.J. and Trewin, N.H.). pdf.
Catalogue of the type and figured material in the Palaeontology Collection, University of Aberdeen, with notes on the H. A. Nicholson collection. Publications of the Department of Geology and Mineralogy, University of Aberdeen 2, 1-28 (Benton, M.J. and Trewin, N.H.). pdf.